4. Diachæa Fries

Sporangia distinct, globose or cylindric, the peridium thin, iridescent, stipitate; the stipe and columella surcharged with lime, white or yellowish, rigid, thick, tapering upward; capillitium of delicate threads free from lime, radiating from various points on the columella, branching and anastomosing as in Comatricha to form a more or less intricate network, the ultimate branchlets supporting the peridial wall.

Rostafinski placed this genus near the Didymieae on account of the calcareous columella and the non-calcareous capillitium. On the other hand the structure of the capillitium and the iridescent simple peridium ally Diachaea to Lamproderma and the Stemoniteae; the only distinction being the calcareous stem. It is simply an intermediate genus to be placed here more conveniently than anywhere else in what is of necessity a linear arrangement.

Key to the Species of Diachæa

A. Stipe and columella white.
a. Sporangium cylindric1. D. leucopodia
b. Sporangium globose.
i. Evidently stalked2. D. splendens
ii. Stalk very short, 5 mm., conic.
O Spores warted3. D. bulbillosa
OO Spores faintly netted4. D. subsessilis
B. Stipe yellowish or orange5. D. thomasii

1. Diachaea leucopodia (Bull.) Rost.

Sporangia rather closely gregarious, metallic blue or purple iridescent, cylindric or ellipsoidal, obtuse, sub-umbilicate below, stipitate; stipe short, much less than one-half the total height, snow-white, tapering upward; hypothallus white, venulose, occurring from stipe to stipe to form an open network over the substratum; columella thick, cylindric, tapering, blunt, terminating below the apex, white; capillitium springing from every part of the columella, of slender threads, brown, flexuous, branching and anastomosing to form an intricate net; spores in mass nearly black, by transmitted light dull violaceous, minutely roughened, 7–9 µ.