Further, but a proportion of the lines of variation is supposed to have been perpetuated, and the extinction of intermediate forms, as already stated, has left isolated groups or species.
The effective cause of these extinctions is stated by Darwin to have been a “natural selection”—a proposition which distinguishes his theory from other development hypotheses, and which is stated in brief by the expression, “the preservation of the fittest.” Its meaning is this: that those characters appearing as results of this spontaneous variation which are little adapted to the conflict for subsistence, with the nature of the supply, or with rivals in its pursuit, dwindle and are sooner or later extirpated; while those which are adapted to their surroundings, and favored in the struggle for means of life and increase, predominate, and ultimately become the centers of new variation. “I am convinced,” says Darwin, “that natural selection has been the main, but not exclusive, means of modification.”
That it has been to a large extent the means of preservation of those structures known as specific, must, I think, be admitted. They are related to their peculiar surroundings very closely, and are therefore more likely to exist under their influence. Thus, if a given genus extends its range over a continent, it is usually found to be represented by peculiar species—one in a maritime division, another in the desert, others in the forest, in the swamp or the elevated areas of the region. The wonderful interdependence shown by Darwin to exist between insects and plants in the fertilization of the latter, or between animals and their food-plants, would almost induce one to believe that it were the true expression of the whole law of development.
But the following are serious objections to its universal application:
First: The characters of the higher groups, from genera up, are rarely of a character to fit their possessors especially for surrounding circumstances; that is, the differences which separate genus from genus, order from order, etc., in the ascending scale of each, do not seem to present a superior adaptation to surrounding circumstances in the higher genus to that seen in the lower genus, etc. Hence, superior adaptation could scarcely have caused their selection above other forms not existing. Or, in other words, the different structures which indicate successional relation, or which measure the steps of progress, seem to be equally well fitted for the same surroundings.
Second: The higher groups, as orders, classes, etc., have been in each geologic period alike distributed over the whole earth, under all the varied circumstances offered by climate and food. Their characters do not seem to have been modified in reference to these. Species, and often genera, are, on the other hand, eminently restricted according to climate, and consequently vegetable and animal food.
The law of development which we seek is indeed not that which preserves the higher forms and rejects the lower after their creation, but that which explains why higher forms were created at all. Why in the results of a creation we see any relation of higher and lower, and not rather a world of distinct types, each perfectly adapted to its situation, but none properly higher than another in an ascending scale, is the primary question. Given the principle of advance, then natural selection has no doubt modified the details; but in the successive advances we can scarcely believe such a principle to be influential. We look rather upon a progress as the result of the expenditure of some force fore-arranged for that end.
It may become, then, a question whether in characters of high grade the habit or use is not rather the result of the acquisition of the structure than the structure the result of the encouragement offered to its assumed beginnings by use, or by liberal nutrition derived from the increasingly superior advantages it offers.
ε. The Physical Origin of Man.
If the hypothesis here maintained be true, man is the descendant of some preëxistent generic type, the which, if it were now living, we would probably call an ape.