The plant gathers these inorganic materials together and makes them up into its own substance. The animal eats the plant and appropriates the nutritious portions to its own sustenance, rejects and gets rid of the useless matters; and, finally, the animal itself dies, and its whole body is decomposed and returned into the inorganic world. There is thus a constant circulation from one to the other, a continual formation of organic life from inorganic matters, and as constant a return of the matter of living bodies to the inorganic world; so that the materials of which our bodies are composed are largely, in all probability, the substances which constituted the matter of long extinct creations, but which have in the interval constituted a part of the inorganic world.
Thus we come to the conclusion, strange at first sight, that the MATTER constituting the living world is identical with that which forms the inorganic world. And not less true is it that, remarkable as are the powers or, in other words, as are the FORCES which are exerted by living beings, yet all these forces are either identical with those which exist in the inorganic world, or they are convertible into them; I mean in just the same sense as the researches of physical philosophers have shown that heat is convertible into electricity, that electricity is convertible into magnetism, magnetism into mechanical force or chemical force, and any one of them with the other, each being measurable in terms of the other,--even so, I say, that great law is applicable to the living world. Consider why is the skeleton of this horse capable of supporting the masses of flesh and the various organs forming the living body, unless it is because of the action of the same forces of cohesion which combines together the particles of matter composing this piece of chalk? What is there in the muscular contractile power of the animal but the force which is expressible, and which is in a certain sense convertible, into the force of gravity which it overcomes? Or, if you go to more hidden processes, in what does the process of digestion differ from those processes which are carried on in the laboratory of the chemist? Even if we take the most recondite and most complex operations of animal life--those of the nervous system, these of late years have been shown to be--I do not say identical in any sense with the electrical processes--but this has been shown, that they are in some way or other associated with them; that is to say, that every amount of nervous action is accompanied by a certain amount of electrical disturbance in the particles of the nerves in which that nervous action is carried on. In this way the nervous action is related to electricity in the same way that heat is related to electricity; and the same sort of argument which demonstrates the two latter to be related to one another shows that the nervous forces are correlated to electricity; for the experiments of M. Dubois Reymond and others have shown that whenever a nerve is in a state of excitement, sending a message to the muscles or conveying an impression to the brain, there is a disturbance of the electrical condition of that nerve which does not exist at other times; and there are a number of other facts and phenomena of that sort; so that we come to the broad conclusion that not only as to living matter itself, but as to the forces that matter exerts, there is a close relationship between the organic and the inorganic world--the difference between them arising from the diverse combination and disposition of identical forces, and not from any primary diversity, so far as we can see.
I said just now that the horse eventually died and became converted into the same inorganic substances from whence all but an inappreciable fraction of its substance demonstrably originated, so that the actual wanderings of matter are as remarkable as the transmigrations of the soul fabled by Indian tradition. But before death has occurred, in the one sex or the other, and in fact in both, certain products or parts of the organism have been set free, certain parts of the organisms of the two sexes have come into contact with one another, and from that conjunction, from that union which then takes place, there results the formation of a new being. At stated times the mare, from a particular part of the interior of her body, called the ovary, gets rid of a minute particle of matter comparable in all essential respects with that which we called a cell a little while since, which cell contains a kind of nucleus in its centre, surrounded by a clear space and by a viscid mass of protein substance (Fig. 2); and though it is different in appearance from the eggs which we are mostly acquainted with, it is really an egg. After a time this minute particle of matter, which may only be a small fraction of a grain in weight, undergoes a series of changes,--wonderful, complex changes. Finally, upon its surface there is fashioned a little elevation, which afterwards becomes divided and marked by a groove. The lateral boundaries of the groove extend upwards and downwards, and at length give rise to a double tube. In the upper and smaller tube the spinal marrow and brain are fashioned; in the lower, the alimentary canal and heart; and at length two pairs of buds shoot out at the sides of the body, and they are the rudiments of the limbs. In fact a true drawing of a section of the embryo in this state would in all essential respects resemble that diagram of a horse reduced to its simplest expression, which I first placed before you (Fig. 1).
Slowly and gradually these changes take place. The whole of the body, at first, can be broken up into "cells," which become in one place metamorphosed into muscle,--in another place into gristle and bone,--in another place into fibrous tissue,--and in another into hair; every part becoming gradually and slowly fashioned, as if there were an artificer at work in each of these complex structures that I have mentioned. This embryo, as it is called, then passes into other conditions. I should tell you that there is a time when the embryos of neither dog, nor horse, nor porpoise, nor monkey, nor man, can be distinguished by any essential feature one from the other; there is a time when they each and all of them resemble this one of the dog. But as development advances, all the parts acquire their speciality, till at length you have the embryo converted into the form of the parent from which it started. So that you see, this living animal, this horse, begins its existence as a minute particle of nitrogenous matter, which, being supplied with nutriment (derived, as I have shown, from the inorganic world), grows up according to the special type and construction of its parents, works and undergoes a constant waste, and that waste is made good by nutriment derived from the inorganic world; the waste given off in this way being directly added to the inorganic world. Eventually the animal itself dies, and, by the process of decomposition, its whole body is returned to those conditions of inorganic matter in which its substance originated.
This, then, is that which is true of every living form, from the lowest plant to the highest animal--to man himself. You might define the life of every one in exactly the same terms as those which I have now used; the difference between the highest and the lowest being simply in the complexity of the developmental changes, the variety of the structural forms, and the diversity of the physiological functions which are exerted by each.
If I were to take an oak tree, as a specimen of the plant world, I should find that it originated in an acorn, which, too, commenced in a cell; the acorn is placed in the ground, and it very speedily begins to absorb the inorganic matters I have named, adds enormously to its bulk, and we can see it, year after year, extending itself upward and downward, attracting and appropriating to itself inorganic materials, which it vivifies, and eventually, as it ripens, gives off its own proper acorns, which again run the same course. But I need not multiply examples,--from the highest to the lowest the essential features of life are the same as I have described in each of these cases.
So much, then, for these particular features of the organic world, which you can understand and comprehend, so long as you confine yourself to one sort of living being, and study that only.
But, as you know, horses are not the only living creatures in the world; and again, horses, like all other animals, have certain limits--are confined to a certain area on the surface of the earth on which we live,--and, as that is the simpler matter, I may take that first. In its wild state, and before the discovery of America, when the natural state of things was interfered with by the Spaniards, the horse was only to be found in parts of the earth which are known to geographers as the Old World; that is to say, you might meet with horses in Europe, Asia, or Africa; but there were none in Australia, and there were none whatsoever in the whole continent of America, from Labrador down to Cape Horn. This is an empirical fact, and it is what is called, stated in the way I have given it you, the "Geographical Distribution" of the horse.
Why horses should be found in Europe, Asia, and Africa, and not in America, is not obvious; the explanation that the conditions of life in America are unfavourable to their existence, and that, therefore, they had not been created there, evidently does not apply; for when the invading Spaniards, or our own yeomen farmers, conveyed horses to these countries for their own use, they were found to thrive well and multiply very rapidly; and many are even now running wild in those countries, and in a perfectly natural condition. Now, suppose we were to do for every animal what we have here done for the horse,--that is, to mark off and distinguish the particular district or region to which each belonged; and supposing we tabulated all these results, that would be called the Geographical Distribution of animals, while a corresponding study of plants would yield as a result the Geographical Distribution of plants.