Contemporaneously with these observations, the indefatigable Ehrenberg had discovered that the "greensands" of the geologist were largely made up of casts of a similar character, and proved the existence of Foraminifera at a very ancient geological epoch, by discovering such casts in a greensand of Lower Silurian age, which occurs near St. Petersburg.

Subsequently, Messrs. Parker and Jones discovered similar casts in process of formation, the original shell not having disappeared, in specimens of the sea-bottom of the Australian seas, brought home by the late Professor Jukes. And the Challenger has observed a deposit of a similar character in the course of the Agulhas current, near the Cape of Good Hope, and in some other localities not yet defined.

It would appear that this infiltration of Foraminifera shells with Glauconite does not take place at great depths, but rather in what may be termed a sublittoral region, ranging from a hundred to three hundred fathoms. It cannot be ascribed to any local cause, for it takes place, not only over large areas in the Gulf of Mexico and the Coast of Florida, but in the South Atlantic and in the Pacific. But what are the conditions which determine its occurrence, and whence the silex, the iron, and the alumina (with perhaps potash and some other ingredients in small quantity) of which the Glauconite is composed, proceed, is a point on which no light has yet been thrown. For the present we must be content with the fact that, in certain areas of the "intermediate zone," greensand is replacing and representing the primitively calcareo- silicious ooze.

The investigation of the deposits which are now being formed in the basin of the Mediterranean, by the late Professor Edward Forbes, by Professor Williamson and more recently by Dr. Carpenter, and a comparison of the results thus obtained with what is known of the surface fauna, have brought to light the remarkable fact, that while the surface and the shallows abound with Foraminifera and other calcareous shelled organisms, the indications of life become scanty at depths beyond 500 or 600 fathoms, while almost all traces of it disappear at greater depths, and at 1,000 to 2,000 fathoms the bottom is covered with a fine clay.

Dr. Carpenter has discussed the significance of this remarkable fact, and he is disposed to attribute the absence of life at great depths, partly to the absence of any circulation of the water of the Mediterranean at such depths, and partly to the exhaustion of the oxygen of the water by the organic matter contained in the fine clay, which he conceives to be formed by the finest particles of the mud brought down by the rivers which flow into the Mediterranean.

However this may be, the explanation thus offered of the presence of the fine mud, and of the absence of organisms which ordinarily live at the bottom, does not account for the absence of the skeletons of the organisms which undoubtedly abound at the surface of the Mediterranean; and it would seem to have no application to the remarkable fact discovered by the Challenger, that in the open Atlantic and Pacific Oceans, in the midst of the great intermediate zone, and thousands of miles away from the embouchure of any river, the sea-bottom, at depths approaching to and beyond 3,000 fathoms, no longer consists of Globigerina ooze, but of an excessively fine red clay.

Professor Thomson gives the following account of this capital discovery:—

"According to our present experience, the deposit of Globigerina ooze is limited to water of a certain depth, the extreme limit of the pure characteristic formation being placed at a depth of somewhere about 2,250 fathoms. Crossing from these shallower regions occupied by the ooze into deeper soundings, we find, universally, that the calcareous formation gradually passes into, and is finally replaced by, an extremely fine pure clay, which occupies, speaking generally, all depths below 2,500 fathoms, and consists almost entirely of a silicate of the red oxide of iron and alumina. The transition is very slow, and extends over several hundred fathoms of increasing depth; the shells gradually lose their sharpness of outline, and assume a kind of 'rotten' look and a brownish colour, and become more and more mixed with a fine amorphous red-brown powder, which increases steadily in proportion until the lime has almost entirely disappeared. This brown matter is in the finest possible state of subdivision, so fine that when, after sifting it to separate any organisms it might contain, we put it into jars to settle, it remained for days in suspension, giving the water very much the appearance and colour of chocolate.

"In indicating the nature of the bottom on the charts, we came, from experience and without any theoretical considerations, to use three terms for soundings in deep water. Two of these, Gl. oz. and r. cl., were very definite, and indicated strongly-marked formations, with apparently but few characters in common; but we frequently got soundings which we could not exactly call 'Globigerina ooze' or 'red clay,' and before we were fully aware of the nature of these, we were in the habit of indicating them as 'grey ooze' (gr. oz.) We now recognise the 'grey ooze' as an intermediate stage between the Globigerina ooze and the red clay; we find that on one side, as it were, of an ideal line, the red clay contains more and more of the material of the calcareous ooze, while on the other, the ooze is mixed with an increasing proportion of 'red clay.'

"Although we have met with the same phenomenon so frequently, that we were at length able to predict the nature of the bottom from the depth of the soundings with absolute certainty for the Atlantic and the Southern Sea, we had, perhaps, the best opportunity of observing it in our first section across the Atlantic, between Teneriffe and St. Thomas. The first four stations on this section, at depths from 1,525 to 2,220 fathoms, show Globigerina ooze. From the last of these, which is about 300 miles from Teneriffe, the depth gradually increases to 2,740 fathoms at 500, and 2,950 fathoms at 750 miles from Teneriffe. The bottom in these two soundings might have been called 'grey ooze,' for although its nature has altered entirely from the Globigerina ooze, the red clay into which it is rapidly passing still contains a considerable admixture of carbonate of lime.