There are two hundred known orders of plants; of these not one is certainly known to exist exclusively in the fossil state. The whole lapse of geological time has as yet yielded not a single new ordinal type of vegetable structure. [4]
The positive change in passing from the recent to the ancient animal world is greater, but still singularly small. No fossil animal is so distinct from those now living as to require to be arranged even in a separate class from those which contain existing forms. It is only when we come to the orders, which may be roughly estimated at about a hundred and thirty, that we meet with fossil animals so distinct from those now living as to require orders for themselves; and these do not amount, on the most liberal estimate, to more than about 10 per cent. of the whole.
There is no certainly known extinct order of Protozoa; there is but one among the Coelenterata—that of the rugose corals; there is none among the Mollusca; there are three, the Cystidea, Blastoidea, and Edrioasterida, among the Echinoderms; and two, the Trilobita and Eurypterida, among the Crustacea; making altogether five for the great sub-kingdom of Annulosa. Among Vertebrates there is no ordinally distinct fossil fish: there is only one extinct order of Amphibia—the Labyrinthodonts; but there are at least four distinct orders of Reptilia, viz. the Ichthyosauria, Plesiosauria, Pterosauria, Dinosauria, and perhaps another or two. There is no known extinct order of Birds, and no certainly known extinct order of Mammals, the ordinal distinctness of the "Toxodontia" being doubtful.
The objection that broad statements of this kind, after all, rest largely on negative evidence is obvious, but it has less force than may at first be supposed; for, as might be expected from the circumstances of the case, we possess more abundant positive evidence regarding Fishes and marine Mollusks than respecting any other forms of animal life; and yet these offer us, through the whole range of geological time, no species ordinally distinct from those now living; while the far less numerous class of Echinoderms presents three; and the Crustacea two, such orders, though none of these come down later than the Paleozoic age. Lastly, the Reptilia present the extraordinary and exceptional phenomenon of as many extinct as existing orders, if not more; the four mentioned maintaining their existence from the Lias to the Chalk inclusive.
Some years ago one of your Secretaries pointed out another kind of positive paleontologic evidence tending towards the same conclusion—afforded by the existence of what he termed "persistent types" of vegetable and of animal life. [5] He stated, on the authority of Dr. Hooker, that there are Carboniferous plants which appear to be generically identical with some now living; that the cone of the Oolitic 'Araucaria' is hardly distinguishable from that of an existing species; that a true 'Pinus' appears in the Purbecks, and a 'Juglans' in the Chalk; while, from the Bagshot Sands, a 'Banksia', the wood of which is not distinguishable from that of species now living in Australia, had been obtained.
Turning to the animal kingdom, he affirmed the tabulate corals of the Silurian rocks to be wonderfully like those which now exist; while even the families of the Aporosa were all represented in the older Mesozoic rocks.
Among the Molluska similar facts were adduced. Let it be borne in mind that 'Avicula', 'Mytails', 'Chiton', 'Natica', 'Patella', 'Trochus', 'Discina', 'Orbicula', 'Lingula', 'Rhynchonella', and 'Nautilus', all of which are existing 'genera', are given without a doubt as Silurian in the last edition of 'Siluria'; while the highest forms of the highest Cephalopods are represented in the Lias by a genus, 'Belemnoteuthis', which presents the closest relation to the existing 'Loligo'.
The two highest groups of the Annulosa, the Insecta and the Arachnida, are represented in the Coal, either by existing genera, or by forms differing from existing genera in quite minor peculiarities.
Turning to the Vertebrata, the only Paleozoic Elasmobranch Fish of which we have any complete knowledge is the Devonian and Carboniferous 'Pleuracanthus', which differs no more from existing Sharks than these do from one another.
Again, vast as is the number of undoubtedly Ganoid fossil Fishes, and great as is their range in time, a large mass of evidence has recently been adduced to show that almost all those respecting which we possess sufficient information, are referable to the same sub-ordinal groups as the existing 'Lepidosteus', 'Polypterus', and Sturgeon; and that a singular relation obtains between the older and the younger Fishes; the former, the Devonian Ganoids, being almost all members of the same sub-order as 'Polypterus', while the Mesozoic Ganoids are almost all similarly allied to 'Lepidosteus'. [6]