Lectures on Evolution / Essay #3 from "Science and Hebrew Tradition"

by Thomas Henry Huxley

You see the singular series of footmarks, drawn of its natural size in the large diagram hanging up here (Fig. 2), which I owe to the kindness of my friend Professor Marsh, with whom I had the opportunity recently of visiting the precise locality in Massachusetts in which these tracks occur. I am, therefore, able to give you my own testimony, if needed, that the diagram accurately represents what we saw. The valley of the Connecticut is classical ground for the geologist. It contains great beds of sandstone, covering many square miles, which have evidently formed a part of an ancient sea-shore, or, it may be, lake-shore. For a certain period of time after their deposition, these beds have remained sufficiently soft to receive the impressions of the feet of whatever animals walked over them, and to preserve them afterwards, in exactly the same way as such impressions are at this hour preserved on the shores of the Bay of Fundy and elsewhere. The diagram represents the track of some gigantic animal, which walked on its hind legs. You see the series of marks made alternately by the right and by the left foot; so that, from one impression to the other of the three-toed foot on the same side, is one stride, and that stride, as we measured it, is six feet nine inches. I leave you, therefore, to form an impression of the magnitude of the creature which, as it walked along the ancient shore, made these impressions.

Fig. 2.—Tracks of Brontozoum.

Of such impressions there are untold thousands upon these sandstones. Fifty or sixty different kinds have been discovered, and they cover vast areas. But, up to this present time, not a bone, not a fragment, of any one of the animals which left these great footmarks has been found; in fact, the only animal remains which have been met with in all these deposits, from the time of their discovery to the present day—though they have been carefully hunted over—is a fragmentary skeleton of one of the smaller forms. What has become of the bones of all these animals? You see we are not dealing with little creatures, but with animals that make a step of six feet nine inches; and their remains must have been left somewhere. The probability is, that they have been dissolved away, and completely lost.

I have had occasion to work out the nature of fossil remains, of which there was nothing left except casts of the bones, the solid material of the skeleton having been dissolved out by percolating water. It was a chance, in this case, that the sandstone happened to be of such a constitution as to set, and to allow the bones to be afterward dissolved out, leaving cavities of the exact shape of the bones. Had that constitution been other than what it was, the bones would have been dissolved, the layers of sandstone would have fallen together into one mass, and not the slightest indication that the animal had existed would have been discoverable.

I know of no more striking evidence than these facts afford, of the caution which should be used in drawing the conclusion, from the absence of organic remains in a deposit, that animals or plants did not exist at the time it was formed. I believe that, with a right understanding of the doctrine of evolution on the one hand, and a just estimation of the importance of the imperfection of the geological record on the other, all difficulty is removed from the kind of evidence to which I have adverted; and that we are justified in believing that all such cases are examples of what I have designated negative or indifferent evidence—that is to say, they in no way directly advance the hypothesis of evolution, but they are not to be regarded as obstacles in the way of our belief in that doctrine.

I now pass on to the consideration of those cases which, for reasons which I will point out to you by and by, are not to be regarded as demonstrative of the truth of evolution, but which are such as must exist if evolution be true, and which therefore are, upon the whole, evidence in favour of the doctrine. If the doctrine of evolution be true, it follows, that, however diverse the different groups of animals and of plants may be, they must all, at one time or other, have been connected by gradational forms; so that, from the highest animals, whatever they may be, down to the lowest speck of protoplasmic matter in which life can be manifested, a series of gradations, leading from one end of the series to the other, either exists or has existed. Undoubtedly that is a necessary postulate of the doctrine of evolution. But when we look upon living Nature as it is, we find a totally different state of things. We find that animals and plants fall into groups, the different members of which are pretty closely allied together, but which are separated by definite, larger or smaller, breaks, from other groups. In other words, no intermediate forms which bridge over these gaps or intervals are, at present, to be met with.

To illustrate what I mean: Let me call your attention to those vertebrate animals which are most familiar to you, such as mammals, birds, and reptiles. At the present day, these groups of animals are perfectly well-defined from one another. We know of no animal now living which, in any sense, is intermediate between the mammal and the bird, or between the bird and the reptile; but, on the contrary, there are many very distinct anatomical peculiarities, well-defined marks, by which the mammal is separated from the bird, and the bird from the reptile. The distinctions are obvious and striking if you compare the definitions of these great groups as they now exist.

The same may be said of many of the subordinate groups, or orders, into which these great classes are divided. At the present time, for example, there are numerous forms of non-ruminant pachyderms, or what we may call broadly, the pig tribe, and many varieties of ruminants. These latter have their definite characteristics, and the former have their distinguishing peculiarities. But there is nothing that fills up the gap between the ruminants and the pig tribe. The two are distinct. Such also is the case in respect of the minor groups of the class of reptiles. The existing fauna shows us crocodiles, lizards, snakes, and tortoises; but no connecting link between the crocodile and lizard, nor between the lizard and snake, nor between the snake and the crocodile, nor between any two of these groups. They are separated by absolute breaks. If, then, it could be shown that this state of things had always existed, the fact would be fatal to the doctrine of evolution. If the intermediate gradations, which the doctrine of evolution requires to have existed between these groups, are not to be found anywhere in the records of the past history of the globe, their absence is a strong and weighty negative argument against evolution; while, on the other hand, if such intermediate forms are to be found, that is so much to the good of evolution; although, for reasons which I will lay before you by and by, we must be cautious in our estimate of the evidential cogency of facts of this kind.

It is a very remarkable circumstance that, from the commencement of the serious study of fossil remains, in fact, from the time when Cuvier began his brilliant researches upon those found in the quarries of Montmartre, palaeontology has shown what she was going to do in this matter, and what kind of evidence it lay in her power to produce.

I said just now that, in the existing Fauna, the group of pig-like animals and the group of ruminants are entirely distinct; but one of the first of Cuvier's discoveries was an animal which he called the Anoplotherium, and which proved to be, in a great many important respects, intermediate in character between the pigs, on the one hand, and the ruminants on the other. Thus, research into the history of the past did, to a certain extent, tend to fill up the breach between the group of ruminants and the group of pigs. Another remarkable animal restored by the great French palaeontologist, the Palaeotherium, similarly tended to connect together animals to all appearance so different as the rhinoceros, the horse, and the tapir. Subsequent research has brought to light multitudes of facts of the same order; and at the present day, the investigations of such anatomists as Rutimeyer and Gaudry have tended to fill up, more and more, the gaps in our existing series of mammals, and to connect groups formerly thought to be distinct.