The family of the Northern crayfishes is termed Potamobiidæ; that of the Southern crayfishes, Parastacidæ. But these two families have in common all those structural characters which are special to neither; and, carrying out the metaphorical nomenclature of the zoologist a stage further, we may say that the two form a Tribe—the definition of which describes the plan which is common to both families.
FIG. 66.—Diagram of the morphological relations of the Astacina.
It may conduce to intelligibility if these results are put into a graphic form. In fig. [66], A. is a diagram representing the plan of an animal in which all the externally visible parts which are found, more or less modified, in the natural objects which we call individual crayfishes are roughly sketched. It represents the plan of the tribe. B. is a diagram exhibiting such a modification of A. as converts it into the plan common to the whole family of the Parastacidæ. C. stands in the same relation to the Potamobiidæ. If the scheme were thoroughly worked out, diagrams representing the peculiarities of {254} form which characterize each of the genera and species, would appear in the place of the names of the former, or of the circles which represent the latter. All these figures would represent abstractions—mental images which have no existence outside the mind. Actual facts would begin with drawings of individual animals, which we may suppose to occupy the place of the dots above the upper line in the diagram.
That all crayfishes may be regarded as modifications of the common plan A, is not an hypothesis, but a generalization obtained by comparing together the observations made upon the structure of individual crayfishes. It is simply a graphic method of representing the facts which are commonly stated in the form of a definition of the tribe of crayfishes, or Astacina.
This definition runs as follows:—
Multicellular animals provided with an alimentary canal and with a chitinous cuticular exoskeleton; with a ganglionated central nervous system traversed by the œsophagus; possessing a heart and branchial respiratory organs.
The body is bilaterally symmetrical, and consists of twenty metameres (or somites and their appendages), of which six are associated into a head, eight into a thorax, and six into an abdomen. A telson is attached to the last abdominal somite.
The somites of the abdominal region are all free, those of the head and thorax, except the hindermost, which is {255} partially free, are united into a cephalothorax, the tergal wall of which has the form of a continuous carapace. The carapace is produced in front into a rostrum, at the sides into branchiostegites.
The eyes are placed at the ends of movable stalks. The antennules are terminated by two filaments. The exopodite of the antenna has the form of a mobile scale. The mandible has a palp. The first and second maxillæ are foliaceous; the second being provided with a large scaphognathite. There are three pairs of maxillipedes, and the endopodites of the third pair are narrow and elongated. The next pair of thoracic appendages is much larger than the rest, and is chelate, as are the two following pairs, which are slender ambulatory limbs. The hindmost two pairs of thoracic appendages are ambulatory limbs, like the foregoing, but not chelate. The abdominal appendages are small swimmerets, except the sixth pair, which are very large, and have the exopodite divided by a transverse joint.