Disregarding the eye-color, the following is a summary of the preceding results: wild-type ♀, 1,001; wild-type ♂, 437; shifted ♂, 328. The result shows that shifted is a sex-linked recessive. The data of table 46 show that the locus of shifted lies about 15 units on one side or the other of vermilion, which from the calculated position of vermilion at 33 would give a position for shifted at either 18 or 48 from yellow.
Table 46.—P1 shifted vermilion ♂ ♂ × wild ♀ ♀. F1 wild-type ♀ × F1 wild-type ♂ ♂.
| Reference. | Females. | Non-cross-over ♂ ♂. | Cross-over ♂ ♂. | Total ♂ ♂. | Cross- over values. | ||
| Shifted vermilion. | Wild- type. | Shifted. | Vermilion. | ||||
| 13 | 345 | 79 | 115 | 8 | 25 | 227 | 15 |
| 29 | 68 | 20 | 32 | 3 | 4 | 59 | 12 |
| 30 | 191 | 37 | 54 | 5 | 13 | 109 | 17 |
| 31 | 151 | 41 | 65 | 17 | 13 | 136 | 22 |
| 33 | 133 | 49 | 40 | 4 | 6 | 99 | 10 |
| 34 | 113 | 56 | 59 | 9 | 11 | 135 | 15 |
| Total. | 1,001 | 282 | 365 | 46 | 72 | 765 | 15 |
LINKAGE OF SHIFTED, VERMILION, AND BAR.
In order to determine on which side of vermilion shifted lies, a shifted vermilion (not-bar) female was crossed to a (not-shifted red) bar male. Three factors are involved, of which one, bar, is dominant. The shifted vermilion (not-bar) stock is a triple recessive, and a three-point back-cross was therefore possible. The daughters were bar and the sons were shifted vermilion (the triple recessive). Inbred these gave the results shown in table 46. The smallest classes (double cross-overs) are shifted and vermilion bar, which places shifted to the left of vermilion at approximately 17.8 units from yellow.
Table 47.—P1 shifted vermilion ♀ × bar ♂ ♂. F1 bar ♀ × F1 shifted vermillion ♂ ♂.
| Reference. |
|
|
|
| Total. | Cross-over values. | ||||||
| Shifted ver- milion. | Bar. | Shifted bar. | Ver- milion | Shifted ver- milion bar. | Wild- type. | Shifted. | Ver- milion bar. | Shifted ver- milion. | Ver- milion bar. | Shifted bar. | ||
| 65 | 56 | 108 | 15 | 20 | 8 | 33 | 1 | 1 | 242 | 15 | 18 | 31 |
The stock of shifted has been thrown away, since too great difficulty was encountered in maintaining it, because, apparently, of sterility in the females.
LETHALS SA AND SB.