[41] Insecta Maderensia, p. 36.
[42] Id. p. 310.
[43] Insecta Maderensia, p. 452.
[44] Insecta Maderensia, p. 11.
[45] Religion of Nature Delineated, p. 84.
CHAPTER IV.
ORGANS AND CHARACTERS OF VARIATION.
Having in the preceding chapter briefly alluded to some of the principal causes by which the outward aspect of the insect tribes would seem to be in a large measure (though within definite specific limits) regulated, it may perhaps be desirable to gather into a small compass, from those remarks, what the chief organs and characters are which appear to be more peculiarly beneath the control of the various influences which we have been just discussing. To imagine that when an insect has become much altered in its general contour, all the parts of which it is composed are equally affected, is contrary to experience; since observation warns us that there are but few actual members which are capable of change,—whilst even the external features, or secondary diagnostics, are only interfered with according to a fixed law, the workings of which are necessarily modified, in proportion as the constitutions of the several animals are differently organized and acted upon.
As regards positive structure, indeed, we can have but few observations to communicate,—seeing that the limbs and appendages themselves are usually of so constant a nature, that disturbing agencies have little or no power to divert them from their typical states. Still, there are occasional facts on record, which would tend to prove that even these are not altogether exempt from the deranging force of certain contingencies from without: the number of the antennal joints, for instance, in the tribes where those organs are multiarticulate, is said to vary; but how far this may be dependent on physical influences, I am not in a position to decide. The connateness of the elytra, again, is a character which we may at any rate define as sub-structural; and this I have myself noticed, at times, to fluctuate, according to the circumstances and conditions of the respective localities in which the particular species obtain. Such is eminently the case with the universal Harpalus (the H. vividus, Dej.) of the Madeiran Group. Speaking of this peculiarity, in my volume on the Coleoptera of those islands, I made the following remarks: "But perhaps its most singular character, and in which it differs from every other Harpalus with which I am acquainted, consists in the tendency of its elytra to become united or soldered together. I say 'the tendency,' because it is not always the case that they are joined (which, since the law exists at all, is perhaps the more remarkable), although in most instances, especially in localities much exposed and but slightly elevated above the sea-shore, they are. I have examples, however, from the upper as well as the lower regions, in which both states are represented; and others again in which the elytra are only partially connected, being free at the apex though firmly attached towards the scutellum. In every instance, however, even where they are united throughout their entire length, a little force will succeed in separating them, showing their structure, as I have indicated in the diagnosis, to be sub-connate rather than connate. But that it does require force to effect the disjunction, when they are really in the condition described, is proved to a demonstration to any one who has seen the remains of the insect beneath the slabs of stone on many of the small adjacent islands where it most abounds, or drifting about over the surface of the rocks,—under which circumstances I have observed them in immense numbers, apparently the accumulation of two or three generations, which the violence of the elements had not been able to sever. It is rare in the sylvan districts to find them joined; nevertheless such is sometimes the case,—thus proving that the peculiarity is not actually essential, but merely one which it is the tendency of the species to assume, and which is more developed in some specimens, and under certain conditions, than in others.[46]"