Because of these points of superiority in the Old World grape, since Valk, Allen and Rogers showed the way, American grape-breeders have sought to unite by hybridization the good characters of the Old World grape with those of the American. Nearly half of the 2000 grapes cultivated in eastern America have more or less European blood in them. Yet, despite the efforts of the breeders, few of these hybrids have commercial value. Whether because they are naturally better fixed, or long cultivation has more firmly established them, the vine characters of Vitis vinifera more often appear in varieties arising as primary hybrids between that and the native species, and the weaknesses of the foreign grape, which prevent their cultivation in America, crop out. Hybrids in which the vinifera blood is more attenuated, as secondary or tertiary crosses, give better results.

Several secondary hybrids now rank among the best of the cultivated grapes. Examples are Brighton and Diamond. The first is a cross between Diana-Hamburg, a hybrid of a Vinifera and a Labrusca, crossed in its turn with Concord, a Labrusca; the second is a cross between Iona, also a hybrid between a Vinifera and a Labrusca, crossed with Concord. Both were grown from seed planted by Jacob Moore, Brighton, New York, in 1870. Brighton was the first secondary hybrid to attract the attention of grape-breeders, and its advent marked an important step in breeding grapes.

The signal success achieved by hybridizers of the European grape with native species quickly led to similar amalgamations among American species. Jacob Rommel, of Morrison, Missouri, beginning work about 1860, hybridized Labrusca and Vulpina grapes so successfully that a dozen or more of his varieties are still cultivated. All are characterized by great vigor and productiveness; and, although they lack the qualities which make good table-grapes, they are among the best for wine-making. Rommel has had many followers in hybridizing native species, chief of whom was the late T. V. Munson, [Fig. 50], Denison, Texas, who literally made every combination of grapes possible, grew thousands of seedlings and produced many valuable varieties.

Improvement by selection.

Selection, continued through successive generations, so important in the improvement of field and garden plants, has played but small part in the domestication of the grape. The period between planting and fruiting is so long that progress would be slow indeed were this method relied on. Moreover, selection, as a method in breeding, is possible only when plants are bred pure, and it is the experience of grape-breeders that in pure breeding this fruit loses in vigor and productiveness and that the variations are exceedingly slight and unstable. Many pure-bred grapes have been raised on the grounds of the New York Agricultural Experiment Station under the eyes of the writer, of which very few have surpassed the parent or have shown promise for the practice of selection.

New varieties from sports.

Bud-sports or mutations now and then arise in grapes. But not more than two or three of the 2000 varieties now under cultivation are suspected of having arisen in this way. It is true that mutations seem to occur rather often in grapes, but they are easily confused with variations due to environment and are usually too vague to lay hands on. Until the causes of these mutations are known and until they can be produced and controlled, but little can be hoped for in the amelioration of grapes through mutations.

Hybridizing the Grape

Hybridization has been the chief means of improving the grape. At present, from what is being accomplished by many workers, it looks as if it will long continue to be the best means of improving this fruit. Since the grape-grower must depend on new varieties for progress, as old varieties cannot be changed, it should be the ambition of growers to produce varieties better than those we now have. Many amateur and professional grape-growers in the past have found breeding grapes a pleasing and profitable hobby, so that much knowledge has accumulated in regard to manipulating the plants in hybridization, and the results that follow in the offspring of hybridization.

How to hybridize.