The grape is preeminently a North American plant. The genus Vitis is a large one, from thirty to fifty species being distinguished for the world; more than half of these are found on this continent. But few other plants in America, or in the world, inhabit such varied and such extended areas. In North America wild grapes abound on the warm, dry soils of New Brunswick and New England, about the Great Lakes in Canada and in the United States, and on the fertile river banks and in the valleys, rich woodlands and thickets of the eastern and southern States. They thrive in the dry woods, sandy sea-plains, and reef-keys of the Carolinas, Georgia and Florida where the vines of the Scuppernong often run more than a hundred feet over trees and shrubs, rioting in natural luxuriance. They flourish in the mountains and limestone hills of the Virginias, Tennessee and Kentucky. They are not so common in the West, yet found in almost all parts of Missouri and Arkansas, and from North Dakota through Kansas to southern Texas. Some wild grape is found in each of the Rocky Mountain States on plain or mountain, or in river chasm or dry canon. Several species are found in New Mexico, Arizona and California, where if they did not furnish the Spanish padres of Santa Fe and San Diego with fruit for wine, they suggested to them the planting of the first successful vineyards in the United States.
How did the grape spread from the Carolinas to California and from subtropical Mexico to the barren plains of Central Canada? Why divide into its manifold forms in the distribution? These questions are of practical import to the grape-grower and breeder who seeks to improve this fruit. The knowledge of the distribution and evolution of plants obtained in the last half century is so complete that these questions present few difficulties to the naturalist of today. In answering them no one would now hold that the numerous species and their sub-divisions were created separately for the regions in which they grow. All would take the ground that the different wild forms come from one ancestral species. We can waive the question as to what the original species was and as to where it first grew.
It is certain that grapes have not been distributed over North America by the hand of man. Probably they have been growing in the regions where they are now found since before the migration of the first savages. The agents of distribution have been natural ones, such as animals, birds, and lake and river currents. These have widened the area of a species to limits imposed by the hostile action of other plants and of animals and by geographical and physical conditions. As a species has encroached upon a new region, climate, soil, all of the conditions of environment, and the contest with other living things, have gradually modified its characters until in time it became so changed that it constituted a new species.
This descent from an original species with plants changed by environment has given us, in America, types of the wild grape as widely diverse as the regions they inhabit. The species found in the forests have developed long slender trunks and branches in their struggle to attain sunlight and air. At least two species are dwarf and shrubby, or infrequently climbing, two to six feet high, growing in dry sands, on rocky hills and mountains where roots must cling to rocks and penetrate into interstices. Still another form runs on the ground and over low bushes and is nearly evergreen, but in the herbarium can hardly be distinguished from a grape whose habit of growth is strikingly different. Some are long-lived, growing and bearing fruit for two or more centuries, while others reach no greater age than the ordinary shrub. Some have enormous stems, a foot or more in diameter, gnarled and picturesque and supporting a great canopy of branch and foliage,[38] while others are slender in stem and graceful, almost delicate, in character of vine. Not less remarkable than the differences in structure is the adaptability of the genus and some of the species to varied climatic conditions. Several of the wild grapes develop full size and display natural luxuriance and fruit-bearing qualities only in the Middle States, but may be found on dry, gravelly, wind-swept hills far to the north or in some hot and humid atmosphere of the South, as if to show indifference to wet or dry, heat or cold.
On the other hand there are many strong points of resemblance between the score or more of species. The organs and characters that do not bear the strain of changed environment, nor suffer in the perpetual warfare of nature, are much the same in all of the species of Vitis. Thus the structure of flowers, fruits and seeds is practically identical; all have naked-tipped tendrils; leaves and leaf-buds are very similar; and the various species usually hybridize freely. They are alike in the unlikeness of individual plants in any of the species; that is, all of the individuals of the genus are most variable and seeds taken from the same vine may produce plants quite unlike one another and quite unlike the parent.
These few facts regarding the evolution and distribution of American grapes lead to two important conclusions:
First, the species are so distributed throughout the United States, and individuals of the species grow in such abundance and luxuriance, as to suggest that we shall be able to improve and domesticate some one or more of them for all of the agricultural regions of the country. For it is proved that nearly all of the wild grapes have horticultural possibilities; and experience with many plants teaches that the boundaries of areas inhabited by the wild species of a given region coincide with those suited to the production of the domesticated plant in that region. It is not possible to tell where the grape-growing regions of the future are to be located; for species and individuals of this fruit are so common that no one can say where the grape is most at home in America.
Second, grapes are so variable and plastic in nature that, were it not known from experience, it could be assumed that they would yield readily to improvement. Besides being variable they hybridize freely and thus the plant-breeder can obtain desirable starting points. There are indications that some of the characters of grapes, at least, follow Mendel’s Law, and when once these have been determined, and the more important unit characters segregated and defined, it ought to be possible to combine and rearrange the characters of this fruit with some system and surely with more certainty than in the past.
This brief introduction leads us to the consideration of American grapes as cultivated plants. We have seen that it is an absolute impossibility to grow the Old World grape in eastern America. The fruit-growers in this great region are forced to plant the native grapes if any. It required two hundred years to establish this fact and it is less than a hundred years since grape-growers have generally acknowledged it as a fact. What was known of American grapes during the two hundred years wasted in attempting to grow the foreign Vinifera? And what has been accomplished in a century in ameliorating the native grapes?
The earliest European visitors to the Atlantic seaboard delighted in the wild grapes which they found everywhere and which reminded them of the Old World vineyards. Had they never seen such a fruit, the wild grapes could not long have escaped their attention; for the Indians knew and used them as they did potatoes, corn, and tobacco. In the narratives of the early voyages the grape is often in the lists of the resources and treasures of the new-found continent. Unfortunately it was not considered of great intrinsic value but only suggested to the explorers that the grape of the old home might be grown in the new home. Could a part of the exaggerated esteem given by the early European travelers and home-seekers to sassafras, ginseng and other such plants, have been bestowed upon the wild grapes which over-run the country, viticulture would have taken rank with the tobacco, lumber and the fish industries of the early settlers.