During the past few years there has been a tendency on the part of some of our biologists engaged in experimental work to disparage the studies of the systematists. It must be granted, however, that both lines of work are essential to the sound development of zoological science, for experimental investigations in which the accurate diagnosis of species is ignored always result in confusion.
Ecology.—The marvelous modifications in structure and instincts by which the various animals are adapted to their surroundings now forms a special topic in biological research and one of the most fascinating. The adaptations in habitat, time, behavior, appearance and even in structure are found capable of a certain individual modification when studied experimentally.
Zoogeography.—Closely associated with systematic zoology, and indeed a part of the subject in its broader sense, is the study of the geographical distribution of animal species and larger groups.
Paleontology.—The geological succession of organisms embraces a field where zoologist and geologist meet. The wonderful progress made by American investigators is well described in the preceding chapters on Historical Geology and Vertebrate Paleontology.
Biometry.
Since Darwin’s theory of evolution postulated the origin of new species by means of natural selection, it was obviously necessary in order to apply a critical test to determine the precise limits of a species. It was, therefore, proposed to subject a given species to a strict examination by the application of statistical methods to determine the range of variation of its members and the extent to which the species intergrades with others. Other problems, particularly those concerning heredity, were treated in similar manner. This branch of biological science was particularly developed by the English School, led by Sir Francis Galton, followed by Karl Pearson and William Bateson.
In America the methods of biometry have been utilized extensively by Charles B. Davenport, Raymond Pearl, H. S. Jennings and others in the solution of problems in genetics and evolution. Their work shows the great value of critical statistical analysis in the interpretation of biological data. A thorough training in mathematics is now found to be hardly less important for the biologist than is a knowledge of physics and chemistry, for the science of biometry has become one of the most important adjuncts to the study of genetics.
Comparative Anatomy and Embryology.
Comparative Anatomy.—Upon the foundations laid down by Cuvier a century ago the present elaborate structure of comparative anatomy of animals, both vertebrate and invertebrate, has been developed. Vast as is the present accumulation of facts and theories many important problems still await their solution. Jeffries Wyman was long a leader in this field, where many workers are now engaged.
Embryology.—The embryological studies, so brilliantly begun by Von Baer early in the nineteenth century, are still in progress. They have now been extended to the groups more difficult of investigation and into the earliest stages of fertilization and implantation in the mammals. Artificial cultural methods have yielded important results. Louis and Alexander Agassiz, Mark, Minot, Brooks, Whitman, Conklin and E. B. Wilson have taken prominent parts in this work.