1. That observed and admitted variations spring up in the course of descents from a common progenitor.

2. That many of these variations tend to an improvement upon the parent stock.

3. That, by a continued selection of these improved specimens as the progenitors of future stock, its powers may be unlimitedly increased.

4. And, lastly, that there is in nature a power continually and universally working out this selection, and so fixing and augmenting these improvements.

Mr. Darwin's whole theory rests upon the truth of these propositions and crumbles utterly away if only one of them fail him. These, therefore, we must closely scrutinise. We will begin with the last in our series, both because we think it the newest and the most ingenious part of Mr. Darwin's whole argument, and also because, whilst we absolutely deny the mode in which he seeks to apply the existence of the power to help him in his argument, yet we think that he throws great and very interesting light upon the fact that such self-acting power does actively and continuously work in all creation around us.

Mr. Darwin finds then the disseminating and improving power, which he needs to account for the development of new forms in nature, in the principle of "Natural Selection," which is evolved in the strife for room to live and flourish which is evermore maintained between themselves by all living things. One of the most interesting parts of Mr. Darwin's volume is that in which he establishes this law of natural selection; we say establishes, because—repeating that we differ from him totally in the limits which he would assign to its action—we have no doubt of the existence or of the importance of the law itself.

* * * * *

We come then to these conclusions. All the facts presented to us in the natural world tend to show that none of the variations produced in the fixed forms of animal life, when seen in its most plastic condition under domestication, give any promise of a true transmutation of species; first, from the difficulty of accumulating and fixing variations within the same species; secondly, from the fact that these variations, though most serviceable for man, have no tendency to improve the individual beyond the standard of his own specific type, and so to afford matter, even if they were infinitely produced, for the supposed power of natural selection on which to work; whilst all variations from the mixture of species are barred by the inexorable law of hybrid sterility. Further, the embalmed records of 3,000 years show that there has been no beginning of transmutation in the species of our most familiar domesticated animals; and beyond this, that in the countless tribes of animal life around us, down to its lowest and most variable species, no one has ever discovered a single instance of such transmutation being now in prospect; no new organ has ever been known to be developed—no new natural instinct to be formed—whilst, finally, in the vast museum of departed animal life which the strata of the earth imbed for our examination, whilst they contain far too complete a representation of the past to be set aside as a mere imperfect record, yet afford no one instance of any such change as having ever been in progress, or give us anywhere the missing links of the assumed chain, or the remains which would enable now existing variations, by gradual approximations, to shade off into unity. On what then is the new theory based? We say it with unfeigned regret, in dealing with such a man as Mr. Darwin, on the merest hypothesis, supported by the most unbounded assumptions. These are strong words, but we will give a few instances to prove their truth:—

All physiologists admit that the swim-bladder is homologous or "ideally similar" in position and structure with the lungs of the higher vertebrate animals; hence there seems to me to be no great difficulty in believing that natural selection has actually converted a swim-bladder into a lung, or organ used exclusively for respiration.—p. 191.

I can indeed hardly doubt that all vertebrate animals having true lungs have descended by ordinary generation from the ancient prototype, of which we know nothing, furnished with a floating apparatus or swim-bladder—p. 191.