Harry Marshall Ward, eldest son of Francis Marshall Ward, was born in Hereford, March 21, 1854, but he came of a Lincolnshire stock, settled for some time in Nottingham. From unavoidable causes he left school at 14, but afterwards continued his education by attending evening classes organised under the Science and Art Department. To that Department, he owed indirectly the opportunity of a useful and brilliant career. His means were small, and his earliest aim was to qualify as a science teacher. He was admitted to a course of instruction for teachers in training given by Prof. Huxley in 1874-5. Although he must have derived from it a sound insight into the principles of zoology, the subject does not seem to have had any permanent attraction for him.
In the summer of 1857 Ward came under my hands in a course of instruction in botany which I conducted with Prof. Vines in the Science Schools at South Kensington, and from this time onwards we were in intimate relations to the close of his life. I can best tell the story as it came under my eyes. It contains much that could not easily be dealt with in any other way.
It was soon apparent that we had got hold of a man of exceptional ability. It must be confessed that the atmosphere was stimulating, and the conditions under which the teaching was carried on necessitated its being given at high pressure. I remember that on one occasion Ward fainted at his work, from no other cause, I think, than over-excitement. In the autumn of the same year he went for one session to Owens College, Manchester, with the object of continuing his general education. I learn that he carried off the prizes in every subject that he took up.
In the succeeding year I was glad to avail myself of the assistance of Ward as demonstrator in a subsequent course at South Kensington, which I undertook with Prof. Vines. Later in the year he became a candidate for and secured an open scholarship at Christ's College, where Vines himself was then a Fellow, and went into residence in October, 1876.
Ward took full advantage of his opportunities at Cambridge, and attended the teaching of Sir Michael Foster in physiology and of Prof. F. M. Balfour in comparative anatomy. The sound and fundamental conceptions which he acquired from the former manifestly influenced his work throughout life. He took a first class in botany in the Natural Science Tripos in 1879. His first published paper was the result of work in the same year in the Jodrell Laboratory at Kew. In this, which was published in the Proceedings of the Linnean Society, he seriously criticised and corrected that of Vesque on the embryo-sac of Phanerogams.
As was customary with our young botanists, Ward went to Germany for a short time, for purposes of study and to strengthen his knowledge of the language. He worked at Würzburg with Sachs, whose lectures on the physiology of plants he afterwards translated in 1887. There he continued his study of the embryo-sac in Orchideae, as Sachs subsequently testified, "zu meiner vollsten Zufriedenheit."
Before the end of the year Ward was appointed on the recommendation of Kew to proceed to Ceylon for two years as Government Cryptogamist to investigate the leaf-disease in coffee. The history of this malady is almost unique in vegetable pathology. A native fungus which had eluded scientific observation, and must therefore have maintained an inconspicuous and limited existence on some native host-plant, found a wider opportunity on the Arabian coffee plant and fell upon it as a devastating scourge. It was first detected in 1869 on a single estate; in 1873 there was probably none in the island entirely free from it. Mr (since Sir Daniel) Morris had shown that the plants could be cleansed by dusting them with a mixture of sulphur and lime. But the remedy proved of no avail as the plants speedily became re-infected. Morris had been transferred to another appointment in the West Indies and Ward's duty was to take up the investigation. This he accomplished exhaustively. He showed that the fungus (Hemileia vastatrix) was one of the Uredineae and that infection was produced by the wind-borne uredospores. Had the planters, as in Southern India, left forest belts between their plantations, the spores might have been filtered out and the disease controlled. As it was it spread like an unchecked conflagration. Ward also discovered the teleutospores; nothing has been added to our knowledge of its life-history beyond what he obtained. The result of his investigations was given in three official reports and in papers contributed in 1882 to the Linnean Society and the Quarterly Journal of Microscopical Science. It was no blame to him that his work led to no practical result. The mischief admitted of no remedy. The coffee-planting industry of Ceylon was destroyed and the Oriental Bank succumbed in the general ruin. Leaf disease has now extended to every coffee-growing country in the Old World from Natal to Fiji.
In a tropical country leaves supply a substratum to a little flora of their own, consisting of organisms partly algal, partly fungal, in their affinity. Ward, who had already developed his characteristic habit of never neglecting any point incidental to a research, carefully studied them, in order both to ascertain how far their presence affected the health of the leaf itself and to work out their life-history. The outcome was three important papers. One on Meliola, an obscure genus of tropical epiphyllous fungi, belonging to the Pyrenomycetes, was published in the Philosophical Transactions in 1883. Bornet's classical memoir published in 1851 had been the authority on the subject. Ward was able to fill up "large gaps in the knowledge of important details." Another paper published in the Quarterly Journal of Microscopical Science in 1882 on an Asterina illuminates an allied organism. But the crown of all Ward's Ceylon work was the splendid memoir on a Tropical Epiphyllous Lichen which was published by the Linnean Society in 1883. In this he, I think, cleared up much that was obscure in the Mycoidea parasitica described by D. D. Cunningham. Having myself communicated the paper, I shall always remember the pleasure with which I undertook in Ward's absence to give an account of it. He solved the problem with convincing completeness; he extended Schwendener's lichen theory to a group of obscure epiphyllous organisms of which he afforded, for the first time, a rational explanation. The success with which this was accomplished placed him at once in the first rank of mycological investigators.
De Bary was the leading authority on Uredineae; and in 1882 Ward paid a short visit to him at Strasburg to confer with him on his coffee disease work, the accuracy of which de Bary entirely confirmed. There he made the acquaintance of Elfving and completed his Meliola paper.
The outlook for Ward was now precarious. Fortunately, I found myself sitting next to Sir Henry Roscoe at a Royal Society dinner, and I suggested that Ward, as an old student of Owens College, would be a fitting recipient of a Bishop Berkeley Fellowship for original research. Principal Greenwood recorded the fact that "the very important results already achieved by Mr Ward in Ceylon, in the domain of the higher botany, led the Senate and the Council to make this appointment." In 1883, he was appointed Assistant Lecturer and Demonstrator in Botany, and, on the same testimony, "abundantly justified his election." It was a peculiar pleasure to him to relieve the veteran Professor Williamson by taking entire charge of Vegetable Physiology and Histology. His position was, in the same year, made secure by his election to a Fellowship at Christ's College, and he married the eldest daughter of the late Francis Kingdon, of Exeter, who was a connection of Clifford the mathematician.