Thus I have been led to recognise species as exemplifying the continuous operation of natural law, or secondary cause; and that not only successively but progressively; "from the first embodiment of the vertebrate idea under its old ichthyic vestment until it became arrayed in the glorious garb of the human form."
III.—Extinction—Cataclysmal or Regulated
If the species of palæothere, paloplothere, anchithere, hipparion, and horse be severally deemed due to remotely and successively repeated acts of creation; the successive going out of such species must have been as miraculous as their coming in. Accordingly, in Cuvier's "Discourse on Revolutions of the Earth's Surface" we have a section of "Proofs that these revolutions have been numerous," and another of "Proofs that these revolutions have been sudden." But as the discoveries of palæontologists have supplied the links between the species held to have perished by the cataclysms, so each successive parcel of geological truth has tended to dissipate the belief in the unusually sudden and violent nature of the changes recognisable in the earth's surface. In specially directing my attention to this moot point, whilst engaged in investigations of fossil remains, I was led to recognise one cause of extinction as being due to defeat in the contest which the individual of each species had to maintain against the surrounding agencies which might militate against its existence. This principle has received a large and most instructive accession of illustrations from the labours of Charles Darwin; but he aims to apply it not only to the extinction but to the origin of species.
Although I fail to recognise proof of the latter bearing of the battle of life, the concurrence of so much evidence in favour of extinction by law is, in like measure, corroborative of the truth of the ascription of the origin of species to a secondary cause.
What spectacle can be more beautiful than that of the inhabitants of the calm expanse of water of an atoll encircled by its ring of coral rock! Leaving locomotive frequenters of the calcarious basin out of the question, we may ask, Was direct creation after the dying out of its result as a "rugose coral" repeated to constitute the succeeding and superseding "tabulate coral"? Must we also invoke the miraculous power to initiate every distinct species of both rugosa and tabulata? These grand old groups have had their day and are utterly gone. When we endeavour to conceive or realise such mode of origin, not of them only but of their manifold successors, the miracle, by the very multiplication of its manifestations, becomes incredible—inconsistent with any worthy conception of an all-seeing, all-provident Omnipotence.
Being unable to accept the volitional hypothesis (of Lamarck) or the selective force exerted by outward circumstances (Darwin), I deem an innate tendency to deviate from parental type, operating through periods of adequate duration, to be the most probable way of operation of the secondary law whereby species have been derived one from another.
According to my derivative hypothesis a change takes place first in the structure of the animal, and this, when sufficiently advanced, may lead to modifications of habits. But species owe as little to the accidental concurrence of environing circumstances as kosmos depends upon a fortuitous concourse of atoms. A purposive route of development and change of correlation and inter-dependence, manifesting intelligent will, is as determinable in the succession of races as in the development and organisation of the individual.
Derivation holds that every species changes in time, by virtue of inherent tendencies thereto. Natural selection holds that no such change can take place without the influence of altered external circumstances educing or eliciting such change.
Derivation sees among the effects of the innate tendency to change, irrespective of altered surrounding circumstances, a manifestation of creative power in the variety and beauty of the results; and, in the ultimate forthcoming of a being susceptible of appreciating such beauty, evidence of the preordaining of such relation of power to the appreciation. Natural selection acknowledges that if power or beauty, in itself, should be a purpose in creation, it would be absolutely fatal to it as a hypothesis.
Natural selection sees grandeur in the "view of life, with its several powers, having been originally breathed by the Creator into a few forms or into one." Derivation sees, therein, a narrow invocation of a special miracle and an unworthy limitation of creative power, the grandeur of which is manifested daily, hourly, in calling into life many forms, by conversion of physical and chemical into vital modes of force, under as many diversified conditions of the requisite elements to be so combined.