Fortunately, however, there is a principle in comparative anatomy which lessens this difficulty. Every organised being constitutes a complete and compact system with all its parts in mutual correspondence. None of its parts can be changed without changing other parts, and consequently each part, taken separately, indicates the others.

Thus, if the intestines of an animal are made to digest raw flesh, its jaws must be likewise constructed to devour prey, its claws to seize and tear it, its teeth to rend it, its limbs to overtake it, its organs of sense to discern it afar. Again, in order to enable the jaw to seize with facility, a certain form of condyle is necessary, and the zygomatic arch must be well developed to give attachment to the masseter muscle. Again, the muscles of the neck must be powerful, whence results a special form in the vertebræ and the occiput, where the muscles are attached. Yet again, in order that the claws may be effective, the toe-bones must have a certain form, and must have muscles and tendons distributed in a certain way. In a word, the form of the tooth necessitates the form of the condyle, of the shoulder-blade, and of the claws, of the femur, and of all the other bones, and all the other bones taken separately will give the tooth. In this manner anyone who is scientifically acquainted with the laws of organic economy may from a fragment reconstruct the whole animal. The mark of a cloven hoof is sufficient to tell the form of the teeth and jaws and vertebræ and leg-bones and thigh-bones and pelvis of the animal. The least fragment of bone, the smallest apophysis, has a determinative character in relation to the class, the order, the genus, and species to which it may belong. This is so true that, if we have only a single extremity of bone well preserved, we may, with application and a skilful use of analogy and exact comparison, determine all those points with as much certainty as if we were in possession of the entire animal. By the application of these principles we have identified and classified the fossil remains of more than one hundred and fifty mammalia.

II.—What the Fossils Teach

An examination of the fossils on the lines I have indicated shows that out of one hundred and fifty mammiferous and oviparous quadrupeds, ninety are unknown to present naturalists, and that in the older layers such oviparous quadrupeds as the ichthyosauri and plesiosauri abound. The fossil elephant, the rhinoceros, the hippopotamus, and the mastodons are not found in the more ancient layers. In fact, the species which appear the same as ours are found only in superficial deposits.

Now, it cannot be held that the present races of animals differ from the ancient races merely by modifications produced by local circumstances and change of climate—for if species gradually changed, we must find traces of these gradual modifications, and between the palæotheria and the present species we should have discovered some intermediate formation; but to the present time none of these have appeared.

Why have not the bowels of the earth preserved the monuments of so remarkable a genealogy unless it be that the species of former ages were as constant as our own, or at least because the catastrophe that destroyed them had not left them time to give evidence of the changes?

Further, an examination of animals shows that though their superficial characteristics, such as colour and size, are changeable, yet their more radical characteristics do not change. Even the artificial breeding of domestic animals can produce only a limited degree of variation. The maximum variation known at the present time in the animal kingdom is seen in dogs, but in all the varieties the relations of the bones remain the same and the shape of the teeth undergoes no palpable change.

I know that some naturalists rely much on the thousands of ages which they can accumulate with a stroke of the pen; but there is nothing which proves that time will effect any more than climate and a state of domestication. I have endeavoured to collect the most ancient documents of the forms of animals. I have examined the engravings of animals including birds on the numerous columns brought from Egypt to Rome. M. Saint Hilaire collected all the mummies of animals he could obtain in Egypt—cats, ibises, birds of prey, dogs, monkeys, crocodiles, etc.—and we cannot find any more difference between them and those of the present day than between human mummies of that date and skeletons of the present day.

There is nothing, then, in known facts which can support the opinion that the new genera discovered among fossils—the palæotheria, anoplotheria, megalonyces, mastodontes, pterodactyli, ichthyosauri, etc.—could have been the sources of any animals now existing, which would differ only by the influence of time or climate.

As yet no human bones have been discovered in the regular layers of the surface of the earth, so that man probably did not exist in the countries where fossil bones are found at the epoch of the revolutions which buried these bones, for there cannot be assigned any reason why mankind should have escaped such overwhelming catastrophes, or why human remains should not be discovered. Man may have inhabited some confined tract of country which escaped the catastrophe, but his establishment in the countries where the fossil remains of land animals are found—that is to say, in the greatest part of Europe, Asia, and America—is necessarily posterior not only to the revolutions which covered these bones, but even to those which have laid open the strata which envelop them; whence it is clear that we can draw neither from the bones themselves nor from the rocks which cover them any argument in favour of the antiquity of the human species in these different countries. On the contrary, in closely examining what has taken place on the surface of the globe, since it was left dry for the last time, we clearly see that the last revolution, and consequently the establishment of present society, cannot be very ancient. An examination of the amount of alluvial matter deposited by rivers, of the progress of downs, and of other changes on the surface of the earth, informs us clearly that the present state of things did not commence at a very remote period.