How have all the exquisite adaptations of one part of the body to another part, and to the conditions of life, and of one organic being to another being, been perfected? For everywhere we find these beautiful adaptations.
The answer is to be found in the struggle for life. Owing to this struggle, variations, however slight, and from whatever cause proceeding, if they be in any degree profitable to the individuals of a species in their infinitely complex relations to other organic beings and to their physical conditions of life, will tend to the preservation of such individuals, and will generally be inherited by the offspring. The offspring, also, will thus have a better chance of surviving, for, of the many individuals of any species which are periodically born, but a small number can survive. I have called this principle, by which each slight variation, if useful, is preserved, by the term Natural Selection, in order to mark its relation to man's power of selection. But the expression, often used by Mr. Herbert Spencer, of the Survival of the Fittest, is more accurate.
We have seen that man, by selection, can certainly produce great results, and can adapt organic beings to his own uses, through the accumulation of slight but useful variations given to him by the hand of Nature. Natural Selection is a power incessantly ready for action, and is as immeasurably superior to man's feeble efforts as the works of Nature are to those of Art.
All organic beings are exposed to severe competition. Nothing is easier than to admit in words the truth of the universal struggle for life, or more difficult—at least, I have found it so—than constantly to bear this conclusion in mind. Yet, unless it be thoroughly engrained in the mind, the whole economy of Nature, with every fact of distribution, rarity, abundance, extinction, and variation, will be dimly seen or quite misunderstood. We behold the face of Nature bright with gladness; we often see superabundance of food. We do not see, or we forget, that the birds which are idly singing round us mostly live on insects or seeds, and are thus constantly destroying life; or we forget how largely these songsters, or their eggs, or their nestlings, are destroyed by birds or beasts of prey. We do not always bear in mind that, though food may be superabundant, it is not so at all seasons of each recurring year.
A struggle for existence, the term being used in a large, general, and metaphorical sense, inevitably follows from the high rate at which all organic beings tend to increase.
Every being, which during its natural lifetime produces several eggs or seeds, must suffer destruction during some period of its life, and during some season or occasional year; otherwise, on the principle of geometrical increase, its numbers would quickly become so inordinately great that no country could support the product. Hence, as more individuals are produced than can possibly survive, there must in every case be a struggle for existence, either one individual with another of the same species, or with the individuals of distinct species, or with the physical conditions of life. It is the doctrine of Malthus applied with manifold force to the whole animal and vegetable kingdoms; for in this case there can be no artificial increase of food, and no prudential restraint from marriage. Although some species may be now increasing, more or less rapidly, in numbers, all cannot do so, for the world would not hold them.
There is no exception to the rule that every organic being naturally increases at so high a rate that, if not destroyed, the earth would soon be covered by the progeny of a single pair. Even slow-breeding man has doubled in twenty-five years, and at this rate, in less than a thousand years, there would literally not be standing-room for his progeny. Linnæus has calculated that if an annual plant produced only two seeds—and there is no plant so unproductive as this—and their seedlings next year produced two, and so on, then in twenty years there would be a million plants. The elephant is reckoned the slowest breeder of all known animals, and I have taken some pains to estimate its probable minimum rate of natural increase. It will be safest to assume that it begins breeding when thirty years old, and goes on breeding until ninety years old, bringing forth six young in the interval, and surviving till one hundred years old. If this be so, after a period of from 740 to 750 years there would be nearly nineteen million elephants alive, descended from the first pair.
The causes which check the natural tendency of each species to increase are most obscure. Eggs or very young animals seem generally to suffer most, but this is not invariably the case. With plants there is a vast destruction of seeds. The amount of food for each species of course gives the extreme limit to which each can increase; but very frequently it is not the obtaining food, but the serving as prey to other animals, which determines the average number of a species. Climate is important, and periodical seasons of extreme cold or drought seem to be the most effective of all checks.
The relations of all animals and plants to each other in the struggle for existence are most complex, and often unexpected. Battle within battle must be continually recurring with varying success; and yet in the long run the forces are so nicely balanced that the face of Nature remains for long periods of time uniform, though assuredly the merest trifle would give the victory to one organic being over another. Nevertheless, so profound is our ignorance, and so high our presumption, that we marvel when we hear of the extinction of an organic being; and as we do not see the cause, we invoke cataclysms to desolate the world, or invent laws on the duration of the forms of life!