How low in the scale of Nature the law of battle descends I know not. Male alligators have been described as fighting, bellowing, and whirling round, like Indians in a war-dance, for the possession of the females; male salmons have been observed fighting all day long; male stag-beetles sometimes bear wounds from the mandibles of other males; the males of certain other insects have been frequently seen fighting for a particular female who sits by, an apparently unconcerned beholder of the struggle, and then retires with the conqueror. The war is, perhaps, severest between the males of the polygamous animals, and these seem oftenest provided with special weapons. The males of carnivorous animals are already well armed, though to them special means of defence may be given through means of sexual selection, as the mane of the lion and the hooked jaw of the salmon. The shield may be as important for victory as the sword or spear.
Amongst birds, the contest is often of a more peaceful character. All those who have attended to the subject believe that there is the severest rivalry between the males of many species to attract, by singing, the females. The rock-thrush of Guiana, birds of paradise, and some others, congregate; and successive males display with the most elaborate care, and show off in the best manner, their gorgeous plumage; they likewise perform strange antics before the females, which, standing by as spectators, at last choose the most attractive partner.
If man can in a short time give beauty and an elegant carriage to his bantams, according to his standard of beauty, I can see no good reason to doubt that female birds, by selecting, during thousands of generations, the most melodious or beautiful males, according to their standard of beauty, might produce a marked effect.
VI.—The Struggle for Existence
Under domestication we see much variability, caused, or at least excited, by changed conditions of life; but often in so obscure a manner that we are tempted to consider the variations as spontaneous. Variability is governed by many complex laws—by correlated growth, compensation, the increased use and disuse of parts, and the definite action of the surrounding conditions. There is much difficulty in ascertaining how largely our domestic productions have been modified; but we may safely infer that the amount has been large, and that modifications can be inherited for long periods. As long as the conditions of life remain the same, we have reason to believe that a modification, which has already been inherited for many generations, may continue to be inherited for an almost infinite number of generations. On the other hand, we have evidence that variability, when it has once come into play, does not cease under domestication for a very long period; nor do we know that it ever ceases, for new varieties are still occasionally produced by our oldest domesticated productions.
Variability is not actually caused by man; he only unintentionally exposes organic beings to new conditions of life, and then Nature acts on the organisation and causes it to vary. But man can and does select the variations given to him by Nature, and thus accumulates them in any desired manner. He thus adapts animals and plants for his own benefit or pleasure. He may do this methodically, or he may do it unconsciously by preserving the individuals most useful or pleasing to him without an intention of altering the breed.
It is certain that he can influence the character of a breed by selecting, in each successive generation, individual differences so slight as to be inappreciable except by an educated eye. This unconscious process of selection has been the agency in the formation of the most distinct and useful domestic breeds. That many breeds produced by man have to a large extent the character of natural species is shown by the inextricable doubts whether many of them are varieties or aboriginally distinct species.
There is no reason why the principles which have acted so efficiently under domestication should not have acted under Nature. In the survival of favoured individuals and races, during the constantly recurrent struggle for existence, we see a powerful and ever-acting form of selection. The struggle for existence inevitably follows from the high geometrical ratio of increase which is common to all organic beings. This high rate of increase is proved by calculation; by the rapid increase of many animals and plants during a succession of peculiar seasons and when naturalised in new countries. More individuals are born than can possibly survive. A grain in the balance may determine which individuals shall live and which shall die; which variety or species shall increase in number, and which shall decrease, or finally become extinct.
As the individuals of the same species come in all respects into the closest competition with each other, the struggle will generally be most severe between them; it will be almost equally severe between the varieties of the same species, and next in severity between the species of the same genus. On the other hand, the struggle will often be severe between beings remote in the scale of Nature. The slightest advantage in certain individuals, at any age or during any season, over those with which they come into competition, or better adaptation, in however slight a degree, to the surrounding physical conditions, will, in the long run, turn the balance.
With animals having separated sexes, there will be in most cases a struggle between the males for the possession of the females. The most vigorous males, or those which have most successfully struggled with their conditions of life, will generally leave most progeny. But success will often depend on the males having special weapons, or means of defence, or charms; and a slight advantage will lead to victory.