Fauna.—California embraces areas of every life-zone of North America: of the boreal, the Hudsonian and Canadian subzones; of the transition, the humid Pacific subzone; of the upper austral, the arid or upper Sonoran subzone; of the lower austral, the arid or lower Sonoran; of the tropical, the “dilute arid” subzone. As will be inferred from the above account of temperature, summer is longer in the north, and localities in the Valley have more hours of heat than do those of south California. Hence that climatic characteristic of the entire Pacific Coast—already referred to and which is of extreme importance in determining the life-zones of California—the great amount of total annual heat supply at comparatively high latitudes. A low summer temperature enables northern species to push far southward, while the high heat total of the year enables southern species to push far north. The resultant intermingling of forms is very marked and characteristic of the Pacific Coast states. The distribution of life-zones is primarily a matter of altitude and corresponds to that of the isotherms. The mountain goat and mountain sheep live in the Sierran upper-land, though long ago well-nigh exterminated. The Douglas red squirrel is ubiquitous in the Sierran forests and their most conspicuous inhabitant. White-tailed deer and especially black-tails are found on the high Sierra; the mule deer, too, although its habitat is now mainly east of the range, on the plateau, is also met with. Grizzly, black, cinnamon and brown bears are all Californian species once common and to-day rare. When Americans began to rule in California elk and antelope herded in great numbers in the Great Valley; the former may to-day sometimes be seen, possibly, in the northern forests, and the latter occasionally cross into the state from Nevada. The sage-hen is abundant on the eastern flank of the Sierra. Grouse, quail, crows and woodpeckers (Melanerpes formicivorus) furnish species characteristic of the state. There are various species of ground-squirrels and gophers, which are very abundant. Noteworthy in the animal life of the lower Sonoran and tropic region are a variety of snakes and lizards, desert rats and mice; and, among birds, the cactus wren, desert thrasher, desert sparrow, Texas night-hawk, mocking-bird and ground cuckoo or road runner (Geococcyx Californianus). The California vulture, the largest flying bird in North America and fully as large as the Andean condor, is not limited to California but is fairly common there. In the zoology and botany of California as of the rest of the Pacific Coast, the distinctions between the upper austral and humid transition zones are largely obliterated; and as one passes southward into the arid lands, life forms of both these zones intermingle with those of the arid transition.

Fish are abundant. The United States fish commission, and an active state commission established in 1869, have done much to preserve and increase this source of food. In 1904 the yield of the fisheries of the three Pacific Coast states was 168,600,000 lbs, valued at $6,681,000,—nearly half that of the New England states, more than one-third that of the Middle Atlantic states and more than that of the South Atlantic and Gulf states combined. Of the total, California yielded between a quarter and a third. A third of her fish comes from the Sacramento river. Some 230—more or less—marine food fishes are to be found in the market at San Francisco. The exports of fish from that port from 1892-1899 were valued at from $2,000,000 to $2,500,000 annually. Native oysters are small and of peculiar flavour; eastern varieties also are fattened, but not bred in California waters. Shrimp are abundant; the shrimp fishers are Chinese and four-fifths of the catch is exported to China. Sturgeon were once the cheapest fish after salmon; to-day, despite all efforts to increase the supply, they are the dearest. Salmon, once threatened with extinction, have been saved, maintained in good supply, and indeed have probably regained their pristine abundance. Shad and striped bass are both very abundant and cheap. Black bass, flounders, terrapin, sea-turtles, perch, turbot, sole and catfish are also common. Great herds of seals once lay like toll-gatherers off the Golden Gate and other bays of the coast, taking a large share of the salmon and other fish; but they are no longer common. The sea-lions sometimes raid the rivers for 100 m. inland. They have greatly increased since hunting them for their hides and oil ceased to be profitable, and thousands sometimes gather on the Farallones, off the Golden Gate.

Flora.—Inclusiveness of range in the distribution of vegetable life is perhaps more suggestive than the distribution of animal species. The variation is from dwarf mountain pine to giant cactus and dates. The humid transition belt is the habitat of California’s magnificent forests. Nut pine, juniper and true sage-brush (Artemisia tridentata) characterize the upper Sonoran,—although the latter grows equally in the transition zone. Cereals, orchard fruits and alfalfa are of primary importance in the upper and of secondary importance in the lower Sonoran. In the arid portions of this and the tropic areas the indigenous plants are creosote, mesquite and alfileria bushes, desert acacias, paloverdes, alkali-heath, salt grass, agaves, yuccas (especially the Spanish-bayonet and Joshua tree) and cactuses. Among exotics the Australian saltbush spreads successfully over the worst alkali land. The introduction of other exotics into these zones,—made humid by irrigation, which converts them, the one into true austro-riparian the other into true humid tropical,—has revolutionized the agricultural, and indeed the whole, economy of California. At the two ends of Cajon Pass, only four or five kilometres apart, are the two utterly distinct floras of the Mohave desert and the San Bernardino valley. Despite the presence of the pass, plants do not spread, so great is the difference of climatic conditions. On the desert the same plant will vary in different years from 4 in. to 10 ft. in height when equally mature, according to the rainfall and other conditions of growth. Many mature plants are not taller than 0.4 to 0.8 in. The tree yucca often attains a height of 20 to 25 ft., and a diameter of 1.5 ft. About 600 species of plants were catalogued in desert California in 1891 by a government botanical party. The flora of the coast islands of California is very interesting. On Santa Cruz Professor Joseph Le Conte found 248 species, nearly all of which are distinctively Californian, 48 being peculiar to the surrounding islands and 28 peculiar to Southern California. Various other things indicate a separation of the islands from the mainland in quaternary times; since which, owing to the later southward movement on the continent of northern forms in glacial times, there has been a struggle for existence on the mainland from which the islands have largely escaped.

Forests.—The forests and agricultural crops of the state demand particular notice. In 1900 the woodland was estimated by the United States census at 22% of the state’s area, and the total stand at 200,000 million ft. of timber. The variety of forest trees is not great, but some of the California trees are unique, and the forests of the state are, with those of Oregon and Washington, perhaps the most magnificent of the world. At least the coniferous forests which make up nine-tenths of California’s woodland surpass all others known in number of species and in the size and beauty of the trees. Forty-six species occur, namely, 32 species of pitch trees (18 pines), 12 species of the cypresses and their allies (2 sequoia), and 2 species of yews or their allies. Peculiar to California are the two species of sequoia (q.v.),—the redwood (S. sempervirens), and the big-tree (S. gigantea), remnants of an earlier age when they were common in other parts of the world. The redwood grows only in a narrow strip on the Coast Range from Southern Oregon (where there are not more than 1000 acres) down nearly to the Golden Gate, in a habitat of heavy rains and heavy fogs. They cover an area of about 2000 sq. m. almost unmixed with other species. One fine grove stands S. of San Francisco near Santa Cruz. These noble trees attain very often a height of more than 300 ft., frequently of 350 and even more, and a butt diameter of more than 15 to 20 ft., with clean, straight fluted trunks rising 200 ft. below the lowest branches. They grow in a very dense timber stand; single acres have yielded 1,500,000 ft. B.M. of lumber, and single trees have cut as high as 100,000 ft. The total stand in 1900 was estimated by the United States census as 75,000,000,000 ft., and the ordinary stand per acre varies from 25,000 to 150,000 ft., averaging probably 60,000 ft. The redwood is being rapidly used for lumber. There is nowhere any considerable young growth from seed, although this mode of reproduction is not (as often stated) unknown; the tree will reproduce itself more than once from the stump (hence its name). In thirty years a tree has been known to grow to a height of 80 ft. and a diameter of 16 in. The wood contains no pitch and much water, and in a green condition will not burn. To this fact it owes its immunity from the forest fires which wreak frightful havoc among the surrounding forests. As the redwood is limited to the Coast Range, so the big tree is limited wholly to the Sierra Nevada. Unlike the redwood the big tree occurs in scattered groves (ten in all) among other species. Its habitat extends some 200 m., from latitude 36° to 39°, nowhere descending much below an altitude of 5000 ft., nor rising above 8000 ft. The most northerly grove and the nearest to San Francisco is the Calaveras Grove near Stockton; the Mariposa Grove just south of the Yosemite National Park, is a state reservation and easily accessible to tourists. The noblest groves are near Visalia, and are held as a national park. The average height is about 275 ft., and the diameter near the ground 20 ft.; various individuals stand over 300 ft., and a diameter of 25 ft. is not rare. One tree measures 35.7 ft. inside the bark 4 ft. above the ground, 10 ft. at 200 ft. above the ground, and is 325 ft. tall. Specimens have been cut down that were estimated to be 1300 and even 2200 years old; many trees standing are presumably 2500 years old. It is the opinion of John Muir that the big tree would normally live 5000 years or more; that the California groves are still in their prime; that, contrary to general ideas, the big tree was never more widely distributed than now, at least not within the past 8000 or 10,000 years; that it is not a decaying species, but that on the contrary “no tree of all the forest is more enduringly established in concord with climate and soil,” growing like the mountain pine even on granite, and in little danger save from the greed of the lumberman; but other excellent authorities consider it as hardly holding its own, especially in the north. Three main wood belts cover the flanks of the Sierra: the lower or main pine belt, the silver fir belt, and the upper pine belt. The sugar pine, the yellow or silver pine and the Douglas spruce (considerably smaller than in Oregon and Washington), are rivals in stature and nobility, all attaining 200 ft. or more when full grown; and the incense cedar reaches a height of 150 ft. In this belt and the following one of firs the big tree also grows. The white silver fir (abies concolor) and the silver or red fir (ab. magnifica), standing 200 to 250 ft., make up almost wholly the main forest belt from 5000 to 9000 ft. for some 450 m. Above the firs come the tamarack, constituting the bulk of the lower Alpine forest; the hardy long-lived mountain pine; the red cedar or juniper, growing even on the baldest rocks; the beautiful hemlock spruce; the still higher white pine, nut pine, needle pine; and finally, at 10,000 to 12,000 ft., the dwarf pine, which grows in a tangle on the earth over which one walks, and may not show for a century’s growth more than a foot of height or an inch of girth. The Nevada slope of the mountains below 7500 ft. is covered with the nut pine down to the sage plains. Its nuts are gathered in enormous amounts by the Indians for food; and it is estimated that the yearly harvest of these nuts exceeds in bulk that of all the cereals of California (John Muir). On the Sierra the underbrush is characterized by the pungent manzanita, the California buckeye and the chamiso; the last two growing equally abundantly on the Coast Range. The chamiso and the manzanita, with a variety of shrubby oaks and thorny plants, often grow together in a dense and sometimes quite impenetrable undergrowth, forming what is known as “chaparral”; if the chamiso occurs alone the thicket is a “chamisal.” The elm, the hickory, the beech, the chestnut, and many others of the most characteristic and useful trees of the eastern states were originally entirely wanting in California. Oaks are abundant; they are especially characteristic of the Great Valley, where they grow in magnificent groves. Up to 1910 national forest reserves amounted to 27,968,510 acres. In 1909 Congress created a national forest to include the big tree groves in Calaveras and Tuolumne counties. One of the noblest redwood areas (that of Santa Cruz county) is a state reservation (created in 1901). Even within reservations almost all the merchantable timber is owned by private individuals. In addition to native trees many others—especially ornamental species—have been successfully introduced from various parts of the world.

Soil.—Sand and loams in great variety, grading from mere sand to adobe, make up the soils of the state. The plains of the north-east counties are volcanic, and those of the south-east sandy. It is impossible to say with accuracy what part of the state may properly be classed as tillable. The total farm acreage in 1900 was 28,828,951 acres, of which 41.5% were improved; since 1880 the absolute amount of improved land has remained practically constant, despite the extraordinary progress of the state in these years. Much land is too rough, too elevated or too arid ever to be made agriculturally available; but irrigation, and the work of the state and national agricultural bureaus in introducing new plants and promoting scientific farming, have accomplished much that once seemed impossible. The peculiarities of the climate, especially its division into two seasons, make Californian (and Southern Arizona) agriculture very different from that of the rest of the country. During the winter no shelter is necessary for live-stock, nor, during summer, for the grains that are harvested in June and July, and may lie for weeks or months in the field. The mild, wet winter is the season of planting and growth, and so throughout the year there is a succession of crops. The dangers of drought in the long dry seasons particularly increase the uncertainties of agriculture in regions naturally arid. Irrigation was introduced in Southern California before 1780, but its use was desultory and its spread slow till after 1850. In 1900 almost 1,500,000 acres were irrigated—an increase of 46% since 1890. About half of this total was in San Joaquin Valley. California has the greatest area of irrigated land of any state in the Union, and offers the most complete utilization of resources. In the south artesian wells, and in the Great Valley the rivers of the Sierra slope, are the main source of water-supply. On nearly all lands irrigated some crops will grow in ordinary seasons without irrigation, but it is this that makes possible selection of crops; practically indispensable for all field and orchard culture in the south, save for a few moist coastal areas, it everywhere increases the yield of all crops and is practised generally all over the state. Of the acreage devoted to alfalfa in 1899, 76.2% was irrigated; of that devoted to subtropical fruits, 71.7%. Small fruits, orchard fruits, hay, garden products and grains are decreasingly dependent on irrigation; wheat, which was once California’s great staple, is (for good, but not for best results) comparatively independent of it,—hence its early predominance in Californian agriculture, due to this success on arid lands since taken over for more remunerative irrigated crops.

Agriculture.—The spread of irrigation and of intensive cultivation, and the increase of small farms during the last quarter of the 19th century, have made California what it is to-day. Agriculture had its beginning in wheat-raising on great ranches, from 50,000 even to several hundred thousand acres in extent. A few of these, particularly in the Great Valley, are still worked, but only a few. The average size of farms in 1850 (when the large Mexican grants were almost the only farms, and these unbroken) was 4466 acres; in 1860 it was 466.4, and in 1900 only 397.4 acres. Stock ranches, tobacco plantations, and hay and grain farms, average from 800 to 530 acres, and counteract the tendency of dairy farms, beet plantations, orchards, vegetable gardens and nurseries to lower the size of the farm unit still further. The renting of large holdings prevails to a greater extent than in any other state except Texas. From 1880 to 1900 the number of farms above 500 and below 1000 acres doubled; half of the total in 1900 were smaller than 100 acres. The most remunerative and most characteristic farming to-day is diversified and intensive and on small holdings. The essential character of California’s economic life has been determined by the successive predominance of grass, gold, grain and fruits. Omitting the second it may be truly said that the order of agricultural development has been mainly one of blind experiment or fortuitous circumstances. Staple products have changed with increasing knowledge of climatic conditions, of life-zones and of the fitness of crops; first hides and tallow, then wool, wheat, grapes (which in the early eighteen-nineties were the leading fruit), deciduous orchard fruits, and semi-tropical citrus fruits successively. Prunes were introduced in 1854, but their possibilities were only slightly appreciated for some thirty years. Of various other crops much the same is true. Of late years progress has been very intelligent; in earlier years it was gained through a multitude of experiments and failures, and great pecuniary loss, and progress was a testimonial chiefly to courage and perseverance. The possibilities of the lower Sonoran and tropical areas are still imperfectly known. Nature has been niggard of rain but lavish in soil and sun. Irrigation has shown that with water, arid and barren plains, veritable deserts may be made to bloom with immense wealth of semi-tropical fruits; and irrigation in the tropical area along the Colorado river, which is so arid that it naturally bears only desert vegetation, has made it a true humid-tropical region like Southern Florida, growing true tropical fruits.

In 1900 California ranked eleventh among the states in total value of farm property ($796,527,955) and in 1899 fourteenth in the value of farm products ($131,690,606). The growth of the former from 1890 to 1900 was only 2.5%, one of the smallest increases among all the states.

The pastoral period extended from 1769 to 1848. The live-stock industry was introduced by the Franciscans and flourished exceedingly. In 1834, when the missions had already passed their best days, there were some 486,000 cattle, horses, mules and asses on the ranges, and 325,000 small animals, principally sheep. Throughout the pre-American period stock-raising was the leading industry; it built up the prosperity of the missions, largely supported the government and almost exclusively sustained foreign commerce. Hides and tallow were the sum and substance of Californian economy. Horses were slaughtered wholesale at times to make way for cattle on the ranges. There was almost no dairying; olive oil took the place of butter, and wine of milk, at the missions; and in general indeed the Mexicans were content with water. In the development of the state under the American regime the live-stock industry has been subordinate. A fearful drought in 1862-1864 greatly depressed it, and especially discouraged cattle ranching. Sheep then became of primary importance, until the increase of the flocks threatened ranges and forests with destruction. As late as 1876 there were some 7,000,000 sheep, in 1900 only 2,581,000, and in 1906 only 1,750,000. In the total value of all live stock (5,402,297 head) in 1900 ($65,000,000) the rank of the state was 15th in the Union, and in value of dairy products in 1899 (12.84 million dollars) 12th. The live-stock industry showed a tendency to decline after 1890, and the dairy industry also, despite various things—notably irrigation and alfalfa culture—that have favoured them.

Cereals replaced hides and tallow in importance after 1848. Wheat was long California’s greatest crop. Its production steadily increased till about 1884, the production in 1880, the banner year, being more than 54 million bushels (32,537,360 centals). Since 1884 its production has markedly fallen off; in 1905 the wheat crop was 17,542,013 bushels, and in 1906, 26,883,662 bushels (valued at $20,162,746). There has been a general parallelism between the amount of rain and the amount of wheat produced; but as yet irrigation is little used for this crop. In the eighth decade of the 19th century, the value of the wheat product had come to exceed that of the annual output of gold. Barley has always been very important. The acreage given to it in 1899 was one-fourth the total cereal acreage, and San Francisco in 1902-1904 was the shipping point of the larger part of American exported barley, of (roughly) three-quarters in 1902, seven-eighths in 1903 and four-fifths in 1904. In 1906 California produced 38,760,000 bushels of barley, valued at $20,930,400. The great increase in the acreage of barley, which was 22.5% of the country’s barley acreage in 1906, and 24.2% in 1905, is one reason for the decreased production of wheat. The level nature of the great grain farms of the valley led to the utilization of machinery of remarkable character. Combined harvesters (which enter a field of standing grain and leave this grain piled in sacks ready for shipment), steam gang-ploughs, and other farm machinery are of truly extraordinary size and efficiency. In 1899 cereals represented more than a third of the total crop acreage and crop product ($93,641,334) of the state. Wheat and other cereals are in part cut for hay, and the hay crop of 1906 was 1,133,465 tons, valued at $12,751,481. California is one of the leading hop-producing states of the Union, the average annual production since 1901 being more than 10,000,000 lb. The product of sugar beets increased between 1888 and 1902 from 1910 to 73,761 tons (according to the state board of trade), and in 1909 (according to the department of agriculture) it was 882,084 tons, from which 254,544,000 lb of sugar was manufactured. In this industry California in 1909 ranked second to Colorado. Truck gardening for export is an assured industry, especially in the north. Great quantities of vegetables, fresh and canned, are shipped yearly, and the same is true on a far larger scale of fruit. Vegetable exports more than doubled between 1894 and 1903. In 1899 hay and grain represented slightly more than a third of the farm acreage and capital and also of the value of all farm products; live-stock and dairy farms represented slightly more than half the acreage, and slightly under 30% of the capital and produce; fruit farms absorbed 6.2% of the acreage and 27% of the capital, and returned 22.5% of the value of farm produce.

Fruit-growing.—Horticulture is now the principal industry, and in this field California has no rival in the United States, although ranking after Florida in the growth of some tropical or semi-tropical fruits,—pineapples, guava, limes, pomeloes or grape-fruit and Japanese persimmons. In 1899 California’s output of fruit was more than a fifth of that of the whole Union. The supremacy of the state is established in the growth of oranges, lemons, citrons, olives, figs, almonds, Persian (or English) walnuts, plums and prunes, grapes and raisins, nectarines, apricots and pomegranates; it also leads in pears, and peaches, but here its primacy is not so assured. Southern California by no means monopolizes the warm-zone fruits. Oranges, lemons and walnuts come chiefly from that section, but citrus fruits grow splendidly in the Sierra foothills of the Sacramento Valley, and indeed ripen earlier there than in the southern district. Almonds, as well as peaches, pears, plums, cherries and apricots, come mainly from the north. Over half of the prune crop comes from Santa Clara county, and the bulk of the raisin output from Fresno county. Olives thrive as far north as the head of the Great Valley, growing in all the valleys and foothills up to 1500 or 2000 ft. They were introduced by the Franciscans (as were various other subtropical fruits, pears and grapes), but their scientific betterment and commercial importance date from about 1885. They grow very abundantly and of the finest quality; for many years poor methods of preparation prejudiced the market against the Californian product, but this has ceased to be the case. The modern orange industry practically began with the introduction into Southern California in 1873 of two seedless orange trees from Brazil; from their stock have been developed by budding millions of trees bearing a seedless fruit known as the “Washington navel,” which now holds first rank in American markets; other varieties, mainly seedlings, are of great but secondary importance. Shipments continue the year round. There has been more than one horticultural excitement in California, but especially in orange culture, which was for a time almost as epidemic a fever as gold seeking once was. By reason of the co-operative effort demanded for the large problems of irrigation, packing and marketing, the citrus industry has done much for the permanent development of the state, and its extraordinary growth made it, towards the close of the 19th century, the most striking and most potent single influence in the growth of agriculture. State legislation has advanced the fruit interest in all possible ways. Between 1872 and 1903 exports of canned fruits increased from 91 to 94,205 short tons; between 1880 and 1903 the increase of dried fruit exports was from 295 to 149,531 tons; of fresh deciduous fruits, from 2590 to 101,199; of raisins, from 400 to 39,963; of citrus fruits, from 458 to 299,623; of wines and brandies between 1891 and 1903, from 47,651 to 97,332 tons. Of the shipments in 1903 some 44% were from Southern California,—i.e. from the seven southernmost counties.