Luminous organs are described by H. von Wielowiejski (Zeits. f. wissen. Zool. xxxvii., 1882); C. Heinemann (Arch. f. mikr. Anat. xxvii., 1886); and R. Dubois (Bull. soc. zool. France, 1886); and stridulating organs by C. J. Gahan (Trans. Entom. Soc., 1900). See also C. Darwin’s Descent of Man and Selection in Relation to Sex (London, 1871).

Many larvae of Coleoptera are described and beautifully figured by J. C. Schiödte (Naturh. Tidsskr. i.-xiii., 1861-1872). Hypermetamorphosis in the Meloidae is described by G. Newport (Trans. Linn. Soc. xx., xxi., 1851-1853); C. V. Riley (Rep. U.S. Entom. Comm. i., 1878); J. H. Fabre (Ann. Sci. Nat. (4), ix., xix., 1848-1853); H. Beauregard (Les Insectes vésicants, Paris, 1890); and A. Chabaud (Ann. Soc. Ent. France, lx., 1891); in the Bruchidae by Riley (Insect Life, iv., v., 1892-1893); and in the Strepsiptera (Stylopidae) by K. T. E. von Siebold (Arch. f. Naturg. ix., 1843); N. Nassonov (Bull. Univ. Narsovie, 1892); and C. T. Brues (Zool. Jahrb. Anat. xiii., 1903).

For various schemes of classification of the Coleoptera see E. L. Geoffroy (Insectes qui se trouvent aux environs de Paris, Paris, 1762); A. G. Olivier (Coléoptères, Paris, 1789-1808); W. S. MacLeay (Annulosa Javanica, London, 1825); the general works of Westwood and Sharp, mentioned above; M. Gemminger and B. de Harold (Catalogus Coleopterorum, 12 vols., Munich, 1868-1872); T. Lacordaire and F. Chapuis (Genera des Coléoptères, 10 vols., Paris, 1854-1874); J. L. Leconte and G. H. Horn (Classification of Coleoptera of N. America, Washington, Smithsonian Inst., 1883); L. Ganglbauer (Die Käfer von Mitteleuropa, Vienna, 1892, &c.); A. Lameere (Ann. Soc. Ent. Belg. xliv., xlvii., 1900-1903); and H. J. Kolbe (Arch. f. Naturg. lxvii., 1901).

For the British species, W. W. Fowler (Coleoptera of the British Islands, 5 vols., London, 1887-1891) is the standard work; and W. F. Johnson and J. N. Halbert’s “Beetles of Ireland” (Proc. R. Irish Acad., 3, vi., 1902) is valuable faunistically. Among the large number of systematic writers on the order generally, or on special families, may be mentioned D. Sharp, T. V. Wollaston, H. W. Bates, G. C. Champion, E. Reitter, G. C. Crotch, H. S. Gorham, M. Jacoby, L. Fairmaire and C. O. Waterhouse.

(G. H. C.)


[1] Instar is a convenient term suggested by D. Sharp to indicate a stage in the life-history of an insect between two successive castings of the cuticle.


COLEPEPER, JOHN COLEPEPER (or Culpepper), 1st Baron (d. 1660), English politician, was the only son of Sir John Colepeper of Wigsell, Sussex. He began his career in military service abroad, and came first into public notice at home through his knowledge of country affairs, being summoned often before the council board to give evidence on such matters. He was knighted, and was elected member for Kent in the Long Parliament, when he took the popular side, speaking against monopolies on the 9th of November 1640, being entrusted with the impeachment of Sir Robert Berkeley on the 12th of February 1641, supporting Stafford’s attainder, and being appointed to the committee of defence on the 12th of August 1641. He separated, however, from the popular party on the Church question, owing to political rather than religious objections, fearing the effect of the revolutionary changes which were now contemplated. He opposed the London petition for the abolition of episcopacy, the project of religious union with the Scots, and the Root and Branch Bill, and on the 1st of September he moved a resolution in defence of the prayer-book. In the following session he opposed the militia bill and the Grand Remonstrance, and finally on the 2nd of January 1642 he joined the king’s party, taking office as chancellor of the exchequer. He highly disapproved of the attempt upon the five members, which was made without his knowledge, but advised the enterprise against Hull. On the 25th of August 1642 he appeared at the bar of the House of Commons to deliver the king’s final proposals for peace, and was afterwards present at Edgehill, where he took part in Prince Rupert’s charge and opposed the retreat of the king’s forces from the battlefield. In December he was made by Charles master of the rolls. He was a leading member of the Oxford Parliament, and was said, in opposition to the general opinion, to have counselled considerable concessions to secure peace. His influence in military affairs caused him to be much disliked by Prince Rupert and the army, and the general animosity against him was increased by his advancement to the peerage on the 21st of October 1644 by the title of Baron Colepeper of Thoresway in Lincolnshire.

He was despatched with Hyde in charge of the prince of Wales to the West in March 1645, and on the 2nd of March 1646, after Charles’s final defeat, embarked with the prince for Scilly, and thence to France. He strongly advocated the gaining over of the Scots by religious concessions, a policy supported by the queen and Mazarin, but opposed by Hyde and other leading royalists, and constantly urged this course upon the king, at the same time deprecating any yielding on the subject of the militia. He promoted the mission of Sir John Berkeley in 1647 to secure an understanding between Charles and the army. In 1648 he accompanied the prince in his unsuccessful naval expedition, and returned with him to the Hague, where violent altercations broke out among the royalist leaders, Colepeper going so far, on one occasion in the council, as to challenge Prince Rupert, and being himself severely assaulted in the streets by Sir Robert Walsh. He continued after the execution of the king to press the acceptance on Charles II. of the Scottish proposals. He was sent to Russia in 1650, where he obtained a loan of 20,000 roubles from the tsar, and, soon after his return, to Holland, to procure military assistance. By the treaty, agreed to between Cromwell and Mazarin, of August 1654, Colepeper was obliged to leave France, and he appears henceforth to have resided in Flanders. He accompanied Charles II. to the south of France in September 1659, at the time of the treaty of the Pyrenees. At the Restoration he returned to England, but only survived a few weeks, dying on the 11th of June 1660.