We begin with artificial wings. The first properly authenticated account of an artificial wing was given by G.A. Borelli in 1670. This author, distinguished alike as a physiologist, mathematician and mechanician, describes and figures a bird with artificial wings, each of which consists of a rigid rod in front and flexible feathers behind. The wings are represented as striking vertically downwards, as the annexed duplicate of Borelli’s figure shows (fig. 31).

Fig. 31.
Borelli’s bird with artificial wings.
r e, Anterior margin of the right wing, consisting of a rigid rod. o a, Posterior margin of the right wing, consisting of flexible feathers. b c, Anterior; and f, Posterior margins of the left wing same as the right. d, Tail of the bird. r g, d h, Vertical direction of the down stroke of the wing.

Borelli was of opinion that flight resulted from the application of an inclined plane, which beats the air, and which has a wedge action. He, in fact, endeavours to prove that a bird wedges itself forward upon the air by the perpendicular vibration of its wings, the wings during their action forming a wedge, the base of which (c b e) is directed towards the head of the bird, the apex (a f) being directed towards the tail (d). In the 196th proposition of his work (De motu animalium, Leiden, 1685) he states that—

“If the expanded wings of a bird suspended in the air shall strike the undisturbed air beneath it with a motion perpendicular to the horizon, the bird will fly with a transverse motion in a plane parallel with the horizon.” “If,” he adds, “the wings of the bird be expanded, and the under surfaces of the wings be struck by the air ascending perpendicularly to the horizon with such a force as shall prevent the bird gliding downwards (i.e. with a tendency to glide downwards) from falling, it will be urged in a horizontal direction.”

The same argument is restated in different words as under:—“If the air under the wings be struck by the flexible portions of the wings (flabella, literally fly flaps or small fans) with a motion perpendicular to the horizon, the sails (vela) and flexible portions of the wings (flabella) will yield in an upward direction and form a wedge, the point of which is directed towards the tail. Whether, therefore, the air strikes the wings from below, or the wings strike the air from above, the result is the same,—the posterior or flexible margins of the wings yield in an upward direction, and in so doing urge the bird in a horizontal direction.”

There are three points in Borelli’s argument to which it is necessary to draw attention: (1) the direction of the down stroke: it is stated to be vertically downwards; (2) the construction of the anterior margin of the wing: it is stated to consist of a rigid rod; (3) the function delegated to the posterior margin of the wing: it is said to yield in an upward direction during the down stroke.

With regard to the first point. It is incorrect to say the wing strikes vertically downwards, for, as already explained, the body of a flying bird is a body in motion; but as a body in motion tends to fall downwards and forwards, the wing must strike downwards and forwards in order effectually to prevent its fall. Moreover, in point of fact, all natural wings, and all artificial wings constructed on the natural type, invariably strike downwards and forwards.

With regard to the second point, viz. the supposed rigidity of the anterior margin of the wing, it is only necessary to examine the anterior margins of natural wings to be convinced that they are in every case flexible and elastic. Similar remarks apply to properly constructed artificial wings. If the anterior margins of natural and artificial wings were rigid, it would be impossible to make them vibrate smoothly and continuously. This is a matter of experiment. If a rigid rod, or a wing with a rigid anterior margin, be made to vibrate, the vibration is characterized by an unequal jerky motion, at the end of the down and up strokes, which contrasts strangely with the smooth, steady fanning movement peculiar to natural wings.

As to the third point, viz. the upward bending of the posterior margin of the wing during the down stroke, it is necessary to remark that the statement is true if it means a slight upward bending, but that it is untrue if it means an extensive upward bending.

Borelli does not state the amount of upward bending, but one of his followers, E.J. Marey, maintains that during the down stroke the wing yields until its under surface makes a backward angle with the horizon of 45°. Marey further states that during the up stroke the wing yields to a corresponding extent in an opposite direction—the posterior margin of the wing, according to him, passing through an angle of 90°, plus or minus according to circumstances, every time the wing rises and falls.