Hexapoda mostly with wings; the wingless forms clearly degraded or modified. Maxillulae vestigial or absent. No locomotor abdominal appendages (except in certain larvae). Young animals always unlike parents, the wing-rudiments developing beneath the larval cuticle and only appearing in a penultimate pupal instar, which takes no food and is usually passive.

Order: Neuroptera.

Biting mandibles; second maxillae completely fused. Prothorax large and free. Membranous, net-veined wings, those of the two pairs closely alike. Six or eight Malpighian tubes. Cerci absent. Larva campodeiform, usually feeding by suction (exceptionally hypermetamorphic with subsequent eruciform instars). Pupa free.

Includes the alder-flies, ant-lions and lacewing-flies. See [Neuroptera].

Order: Coleoptera.

Biting mandibles; second maxillae very intimately fused. Prothorax large and free. Fore-wings modified into firm elytra, beneath which the membranous hind-wings (when present) can be folded. Cerci absent. Four or six Malpighian tubes. Larva campodeiform or eruciform. Pupa free.

Includes the beetles and the parasitic Stylopidae, often regarded as a distinct order (Strepsiptera). (See [Coleoptera].)

Order: Mecaptera.

Biting mandibles; first maxillae elongate; second maxillae completely fused. Prothorax small. Two pairs of similar, membranous wings, with predominantly longitudinal neuration. Six Malpighian tubes. Larva eruciform. Pupa free. Cerci present.

Includes the single family of Panorpidae (scorpion-flies), often comprised among the Neuroptera.