"It has been noticed," writes our great zoölogist, "that certain types, which are frequently prominent among the representatives of past ages, combine in their structure peculiarities which at later periods are only observed separately in different, distinct types. Sauroid fishes before reptiles, Pterodactyles before birds, Ichthyosauri before dolphins, etc. There are entire families, of nearly every class of animals, which in the state of their perfect development exemplify such prophetic relations…. The sauroid fishes of the past geological ages are an example of this kind. These fishes, which preceded the appearance of reptiles, present a combination of ichthyic and reptilian characters not to be found in the true members of this class, which form its bulk at present. The Pterodactyles, which preceded the class of birds, and the Ichthyosauri, which preceded the Cetaeca, are other examples of such prophetic types."[a]

[Footnote a: Agassiz, Contributions: Essay on Classification, p. 117, where, we may be permitted to note, the word "Crustacea" is by a typographical error printed in place of Cetacea.]

Now these reptile-like fishes, of which gar-pikes are the living representatives, though of earlier appearance, are admittedly of higher rank than common fishes. They dominated until reptiles appeared, when they mostly gave place to—or, as the derivationists will insist, were resolved by divergent variation and natural selection into—common fishes, destitute of reptilian characters, and saurian reptiles, the intermediate grades, which, according to a familiar piscine saying, are "neither fish, flesh, nor good red-herring," being eliminated and extinguished by natural consequence of the struggle for existence which Darwin so aptly portrays. And so, perhaps, of the other prophetic types. Here type and antitype correspond. If these are true prophecies, we need not wonder that some who read them in Agassiz's book will read their fulfilment in Darwin's.

Note also, in tins connection, that, along with a wonderful persistence of type, with change of species, genera, orders, etc., from formation to formation, no species and no higher group which has once unequivocally died out ever afterwards reappears. Why is this, but that the link of generation has been sundered? Why, on the hypothesis of independent originations, were not failing species re-created, either identically or with a difference, in regions eminently adapted to their well-being? To take a striking case. That no part of the world now offers more suitable conditions for wild horses and cattle than the Pampas and other plains of South America is shown by the facility with which they have there run wild and enormously multiplied, since introduced from the Old World not long ago. There was no wild American stock. Yet in the times of the Mastodon and Megatherium, at the dawn of the present period, wild horses and cattle—the former certainly very much like the existing horse—roamed over those plains in abundance. On the principle of original and direct created adaptation of species to climate and other conditions, why were these types not reproduced, when, after the colder intervening era, those regions became again eminently adapted to such animals? Why, but because, by their complete extinction in South America, the line of descent was here utterly broken? Upon the ordinary hypothesis, there is no scientific explanation possible of this series of facts, and of many others like them. Upon the new hypothesis, "the succession of the same types of structure within the same areas during the later geological periods ceases to be mysterious, and is simply explained by inheritance." Their cessation is failure of issue.

Along with these considerations the fact (alluded to on p. 114) should be remembered, that, as a general thing, related species of the present age are geographically associated. The larger part of the plants, and still more of the animals, of each separate country are peculiar to it; and, as most species now flourish over the graves of their by-gone relatives of former ages, so they now dwell among or accessibly near their kindred species.

Here also comes in that general "parallelism between the order of succession of animals and plants in geological times, and the gradation among their living representatives" from low to highly organized, from simple and general to complex and specialized forms; also "the parallelism between the order of succession of animals in geological times—and the changes their living representatives undergo during their embryological growth,"—as if the world were one prolonged gestation. Modern science has much insisted on this parallelism, and to a certain extent is allowed to have made it out. All these things, which conspire to prove that the ancient and the recent forms of life "are somehow intimately connected together in one grand system," equally conspire to suggest that the connection is one similar or analogous to generation. Surely no naturalist can be blamed for entering somewhat confidently upon a field of speculative inquiry which here opens so invitingly; nor need former premature endeavors and failures utterly dishearten him.

All these things, it may naturally be said, go to explain the order, not the mode, of the incoming of species. But they all do tend to bring out the generalization expressed by Mr. Wallace in the formula, that "every species has come into existence coincident both in time and space with preëxisting closely allied species." Not, however, that this is proved even of existing species as a matter of general fact. It is obviously impossible to prove anything of the kind. But we must concede that the known facts strongly suggest such an inference. And since species are only congeries of individuals, and every individual came into existence in consequence of preëxisting individuals of the same sort, so leading up to the individuals with which the species began, and since the only material sequence we know of among plants and animals is that from parent to progeny, the presumption becomes exceedingly strong that the connection of the incoming with the preëxisting species is a genealogical one.

Here, however, all depends upon the probability that Mr. Wallace's inference is really true. Certainly it is not yet generally accepted; but a strong current is setting towards its acceptance.

So long as universal cataclysms were in vogue, and all life upon the earth was thought to have been suddenly destroyed and renewed many times in succession, such a view could not be thought of. So the equivalent view maintained by Agassiz, and formerly, we believe, by D'Orbigny, that, irrespectively of general and sudden catastrophes, or any known adequate physical cause, there has been a total depopulation at the close of each geological period or formation, say forty or fifty times, or more, followed by as many independent great acts of creation, at which alone have species been originated, and at each of which a vegetable and an animal kingdom were produced entire and complete, full-fledged, as flourishing, as wide-spread and populous, as varied and mutually adapted from the beginning as ever afterwards,—such a view, of course, supersedes all material connection between successive species, and removes even the association and geographical range of species entirely out of the domain of physical causes and of natural science. This is the extreme opposite of Wallace's and Darwin's view, and is quite as hypothetical. The nearly universal opinion, if we rightly gather it, manifestly is, that the replacement of the species of successive formations was not complete and simultaneous, but partial and successive; and that along the course of each epoch some species probably were introduced, and some, doubtless, became extinct. If all since the tertiary belongs to our present epoch, this is certainly true of it: if to two or more epochs, then the hypothesis of a total change is not true of them.

Geology makes huge demands upon time; and we regret to find that it has exhausted ours,—that what we meant for the briefest and most general sketch of some geological considerations in favor of Darwin's hypothesis has so extended as to leave no room for considering "the great facts of comparative anatomy and zoölogy" with which Darwin's theory "very well accords," nor for indicating how "it admirably serves for explaining the unity of composition of all organisms, the existence of representative and rudimentary organs, and the natural series which genera and species compose." Suffice it to say that these are the real strongholds of the new system on its theoretical side; that it goes far towards explaining both the physiological and the structural gradations and relations between the two kingdoms, and the arrangement of all their forms in groups subordinate to groups, all within a few great types; that it reads the riddle of abortive organs and of morphological conformity, of which no other theory has ever offered a scientific explanation, and supplies a ground for harmonizing the two fundamental ideas which naturalists and philosophers conceive to have ruled the organic world, though they could not reconcile them, namely: Adaptation to Purpose and the Conditions of Existence, and Unity of Type. To reconcile these two undeniable principles is a capital problem in the philosophy of natural history; and the hypothesis which consistently does so thereby secures a great advantage.