We have found, then, Radiates, Mollusks, and Articulates in plenty; and now what is to be said of Vertebrates in these old times,—of the highest and most important division of the Animal Kingdom, that to which we ourselves belong? They were represented by Fishes alone; and the Fish chapter in the history of the early organic world is a curious, and, as it seems to me, a very significant one. We shall find no perfect specimens; and he would be a daring, not to say a presumptuous thinker, who would venture to reconstruct a fish of the Silurian age from any remains that are left to us. But still we find enough to indicate clearly the style of those old fishes, and to show, by comparison with the living types, to what group of modern times they belong. We should naturally expect to find the Vertebrates introduced in their simplest form; but this is by no means the case: the common fishes, as Cod, Herring, Mackerel, and the like, were unknown in those days.
But there are two groups of so-called fishes, differing from these by some marked features, among which we may find the modern representatives of these earliest Vertebrates. Of these two groups one consists chiefly now of the Gar-Pikes of our Western waters, though the Sturgeons share also in some of their features. In these fishes there is a singular union of reptilian with fish-like characters. The systems of circulation and of respiration in them are more complicated than in the common fishes; the structure of the skull resembles that of the skull in reptiles, and they have other reptilian characters, such as their ability to move the head upon the neck independently of the body, and the connection of the vertebrae by ball-and-socket joint, instead of by inverted cones, as in the ordinary fishes. Their scales are also peculiar, being covered by enamel so hard, that, if struck with steel, they will emit sparks like flint. It is on account of this peculiarity that the whole group has been called Ganoid. Now, though we have not found as yet any complete specimens of Silurian fishes, their disconnected remains are scattered profusely in the early deposits. The scales, parts of the backbone, parts of the skull, the teeth, are found in a tolerable state of preservation; and these indications, fragmentary as they are, give us the clue to the character of the most ancient fishes. A large proportion of them were no doubt Ganoids; for they had the same peculiar articulation of the vertebrae, the flexibility of the neck, and the hard scales so characteristic of our Gar-Pikes.
There is another type of these ancient Vertebrates, which has also its representatives among our modern fishes. These are the Sharks and Skates, or, as the Greeks used to call them, the Selachians,—making a very appropriate distinction between them and common fishes, on account of the difference in the structure of the skeleton. In Selachians the quality of the bones is granular, instead of fibrous, as in fishes; the arches above and below the backbone are formed by flat plates, instead of the spines so characteristic of all the fish proper; and the skull consists of a solid box, instead of being built of overlapping pieces like the true fish-skull. They differ also in their teeth, which, instead of being implanted in the bone by a root, as in fishes, are loosely set in the gum without any connection with the bone, and are movable, being arranged in several rows one behind another, the back rows moving forward to take the place of the front ones when the latter are worn off. They are unlike the common fishes also in having the backbone continued to the very end of the tail, which is cut in uneven lobes, the upper lobe being the longer of the two, while the terminal fin, so constant a feature in fishes, is wanting. The Selachians resemble higher Vertebrate types not only in the small number of their eggs, and in the closer connection of the young with the mother, but also in their embryological development, which has many features in common with that of birds and turtles. Of this group, also, we find numerous remains in the ancient geological deposits; and though we have not the means of distinguishing the species, we have ample evidence for determining the type.
This combination of higher with lower features in the earlier organic forms is very striking, and becomes still more significant when we find that many of the later types recall the more ancient ones. I have called these more comprehensive groups of former times, combining characters of different classes, synthetic or prophetic types; and we might as fitly give the name of retrospective types to many of the later groups, for they recall the past, as the former anticipate the future. And it is not only among the Fishes and the Reptiles that we find these combinations. The most numerous of the ancient Radiates are the Acalephlan Corals, combining, in the Hydroid form, the Polyp-like mode of life, habits, and general appearance with the structure of Acalephs. The Crinoids, with the closed cups in some, and the open, star-like crowns in others, unite features of the present Star-Fishes and Sea-Urchins, and, by their stem attaching them to the ground, include also a Polyp-like character; while the Trilobites, with their uniform rings and their prominent anterior shield, unite characters of Worms and Crustacea.
These early types seem to sketch in broad, general characters the Creative purpose, and to include in the first average expression of the plan all its structural possibilities. The Crinoid forms include the thought of the modern Star-Fishes and Sea-Urchins; the simple chambered shells of the Silurian anticipate the more complicated structure of the later ones; the Trilobites give the most comprehensive expression of the Articulate type; while the early Fishes not only prophesy the Reptiles which are to come, but also hint at Birds and even at Mammalia by their embryonic development and their mode of reproduction.
Looked at from this point of view, the animal world is an intellectual Creation, complete in all its parts, and coherent throughout; and when we find, that, although these ancient types have become obsolete and been replaced by modern ones, yet there are always a few old-fashioned individuals, left behind, as it were, to give the key to the history of their race, as the Gar-Pike, for instance, to explain the ancient Fishes, the Millepore to explain the old Acalephian Corals, the Nautilus to be the modern exponent of the Ammonites and Orthoceratites of past times, we cannot avoid the impression that this Creative work has been intended also to be educational for Man, and to teach him his own relation to the organic world. The embryology of the modern types confirms this idea, for here we find an epitome of their geological history. The embryo of the present Star-Fishes recalls the Crinoids; the embryo of the Crab recalls the Trilobites; the embryo of the Vertebrates, including even that of the higher Mammalia, recalls the ancient Fishes. Does not this fact, that the individual animal in its growth recalls the history of its type, prove that the Creative Thought in its immediate present action embraces all that has gone before, as its first organic expression included all that was to come? The study of Nature in its highest meaning shows us the present doubly rich with all the past, and the past linked and interwoven with the present, not lying divorced and dead behind it.
I have spoken of the Silurian beach as if there were but one, not only because I wished to limit my sketch, and to attempt at least to give it the vividness of a special locality, but also because a single such shore will give us as good an idea of the characteristic fauna of the time as if we drew our material from a wider range. There are, however, a great number of parallel ridges belonging to the Silurian and Devonian periods, running from east to west, not only through the State of New York, but far beyond, through the States of Michigan and Wisconsin into Minnesota; one may follow nine or ten such successive shores in unbroken lines, from the neighborhood of Lake Champlain to the Far West. They have all the irregularities of modern sea-shores, running up to form little bays here, and jutting out in promontories there; and upon each one are found animals of the same kind, but differing in species from those of the preceding.
Although the early geological periods are more legible in North America, because they are exposed over such extensive tracts of land, yet they have been studied in many other parts of the globe. In Norway, in Germany, in France, in Russia, in Siberia, in Kamtchatka, in parts of South America, in short, wherever the civilization of the white race has extended, Silurian deposits have been observed, and everywhere they bear the same testimony to a profuse and varied creation. The earth was teeming then with life as now, and in whatever corner of its surface the geologist finds the old strata, they hold a dead fauna as numerous as that which lives and moves above it. Nor do we find that there was any gradual increase or decrease of any organic forms at the beginning and close of the successive periods. On the contrary, the opening scenes of every chapter in the world's history have been crowded with life, and its last leaves as full and varied as its first.
I think the impression that the faunae of the early geological periods were more scanty than those of later times arises partly from the fact that the present creation is made a standard of comparison for all preceding creations. Of course, the collections of living types in any museum must be more numerous than those of fossil forms, for the simple reason that almost the whole of the present surface of the earth, with the animals and plants inhabiting it, is known to us, whereas the deposits of the Silurian and Devonian periods are exposed to view only over comparatively limited tracts and in disconnected regions. But let us compare a given extent of Silurian or Devonian sea-shore with an equal extent of sea-shore belonging to our own time, and we shall soon be convinced that the one is as populous as the other. On the New-England coast there are about one hundred and fifty different kinds of fishes, in the Gulf of Mexico two hundred and fifty, in the Red Sea about the same. We may allow in present times an average of two hundred or two hundred and fifty different kinds of fishes to an extent of ocean covering about four hundred miles. Now I have made a special study of the Devonian rocks of Northern Europe, in the Baltic and along the shore of the German Ocean. I have found in those deposits alone one hundred and ten kinds of fossil fishes. To judge of the total number of species belonging to those early ages by the number known to exist now is about as reasonable as to infer that because Aristotle, familiar only with the waters of Greece, recorded less than three hundred kinds of fishes in his limited fishing-ground, therefore these were all the fishes then living. The fishing-ground of the geologist in the Silurian and Devonian periods is even more circumscribed than his, and belongs, besides, not to a living, but to a dead world, far more difficult to decipher.
But the sciences of Geology and Palaeontology are making such rapid progress, now that they go hand in hand, that our familiarity with past creations is daily increasing. We know already that extinct animals exist all over the world: heaped together under the snows of Siberia,—lying thick beneath the Indian soil,—found wherever English settlers till the ground or work the mines of Australia,—figured in the old Encyclopaedias of China, where the Chinese philosophers have drawn them with the accuracy of their nation,—built into the most beautiful temples of classic lands, for even the stones of the Parthenon are full of the fragments of these old fossils, and if any chance had directed the attention of Aristotle towards them, the science of Palaeontology would not have waited for its founder till Cuvier was born,—in short, in every corner of the earth where the investigations of civilized men have penetrated, from the Arctic to Patagonia and the Cape of Good Hope, these relics tell us of successive populations lying far behind our own, and belonging to distinct periods of the world's history.