Such a critical revision of the fishes of Lake Superior, and the other great Canadian lakes, was the first necessary step in the investigation I am tracing, in order to ascertain the natural primitive relations between them and the region which they inhabit. Before drawing the conclusions which follow directly from these facts, I should introduce a similar list of the fishes living in similar latitudes, or under similar circumstances, in other parts of the world; and more particularly of the species of Northern Europe. But such a list, to be of any use, should be throughout based upon a critical comparative investigation of all the species of that continent, which would lead to too great a digression. The comparison of the fresh-water fishes of Europe, which correspond to those of North America, has been carried so far, that I feel justified in assuming, what is really the fact, that all the species of North America, without a single exception, differ from those of Europe, if we limit ourselves strictly to fishes which are exclusively the inhabitants of fresh water.
I am well aware that the salmon which runs up the rivers of Northern and Central Europe, also occurs on the eastern shores of the northern part of North America, and runs up the rivers emptying into the Atlantic. But this fish is one of the marine arctic fishes, which migrates with many others, annually further south, and which migratory species is common to both continents. Those species, however, which never leave the fresh waters, are, without exception, different on the two continents. Again, on each of the continents, they differ in various latitudes; some, however, taking a wider range than others in their natural geographical distribution.
The fresh-water fishes of North America, which form a part of its temperate fauna, extend over very considerable ground; for there is no reason to subdivide into distinct faunæ the extensive tracts of lands between the arctics and the Middle States of the Union. We notice over these, considerable uniformity in the character of the fresh-water fishes. Nevertheless, a minute investigation of all their species has shewn that Lake Superior proper, and the fresh waters north of it, constitute in many respects a special zoological district, sufficiently different from that of the lower lakes and the northern United States, to form a natural division in the great fauna of the fresh-water fishes of the temperate zone of this continent.
We have shewn that there are types, occurring in all the lower lakes, which never occur in Lake Superior and northwards, and that most of the species found in Lake Superior are peculiar to it; the Salmonidæ only taking a wider range, and some of them covering almost the whole extent of that fauna, while others appear circumscribed within very narrow limits.
Now, such differences in the range which the isolated species take in the faunæ, is a universal character of the distribution of animals; some species of certain families covering, without distinction, extensive grounds, which are occupied by several species of other families, limited to particular districts of the same zone.
But after making due allowance for such variations, and taking a general view of the subject, we arrive, nevertheless, at this conclusion; that all the fresh-water fishes of the district under examination are peculiar to that district, and occur nowhere else in any other part of the world.
They have their analogues in other continents, but nowhere beyond the limits of the American continent do we find any fishes identical with those of the district, the fauna of which we have been recently surveying. The lamprey eels of the lake district have very close representatives in Europe, but they cannot be identified. The sturgeons of this continent are neither identical with those of Europe nor with those of Asia. The cat-fishes are equally different. We find a similar analogy and similar differences between the perches, pickerels, eelpouts, salmons, and carps. In all the families which occur throughout the temperate zone, there are near relatives on the two continents, but they do not belong to the same stock. And in addition to these, there are also types which are either entirely peculiar to the American continent, such as Lepidosteus and Percopsis, or belong to genera which have not simultaneous representatives in the two worlds, and are therefore more or less remote from those which have such close analogues. The family of Percoids, for instance, has several genera in Europe, which have no representatives in America; and several genera in America which have no representatives in Europe, besides genera which are represented on both continents, though by representatives specifically distinct.
Such facts have an important bearing upon the history of creation; and it would be very unphilosophical to adhere to any view respecting its plan, which would not embrace these facts, and grant them their full meaning. If we face the fundamental question which is at the bottom of this particular distribution of animals, and ask ourselves, where have all these fishes been created, there can be but one answer given which will not be in conflict and direct contradiction with the facts themselves, and the laws that regulate animal life. The fishes, and all other fresh-water animals of the region of the great lakes, must have been created where they live. They are circumscribed within boundaries over which they cannot pass, and to which there is no natural access from other quarters. There is no trace of their having extended further in their geographical distribution at any former period, nor of their having been limited within narrower boundaries.
It cannot be rational to suppose that they were created in some other part of the world, and were transferred to this continent, to die away in the region where they are supposed to have originated, and to multiply in the region where they are found. There is no reason why we should not take the present evidence in their distribution as the natural fact respecting their origin, and that they are, and were from the beginning, best suited for the country where they are now found.
Moreover, they bear to the species which inhabit similar regions, and live under similar circumstances in Europe and Asia, and the Pacific side of this continent, such relations, that they appear to the philosophical observer as belonging to a plan which has been carried out in its details with reference to the general arrangement. The species of Europe, Asia, and the Pacific side of this continent, correspond in their general combination to the species of the eastern and northern parts of the American continent, all over which the same general types are extended. They correspond to each other on the whole, but differ as to species.