Elementary Zoology, Second Edition

by Vernon L. Kellogg

The structure and life-history of an example of the polyps (the Fresh-water Hydra, Hydra sp.) has been studied in Chapters [X] and [XI].

OTHER POLYPS, SEA-ANEMONES, CORALS, AND JELLYFISHES

Technical Note.—The teacher should have, if possible, several pieces of coral and a few specimens of Cœlenterates in alcohol or formalin, which will show the external character, at least, of these animals (see account of laboratory equipment, p. [450]). If the school is on the coast, the pupils should be shown the sea-anemones of the tide-pools.

The animals which are included in the branch Cœlenterata are, at least in living condition, unfamiliar to most of us. Like the sponges, they are almost all inhabitants of the ocean; a few, like Hydra, live in fresh water. Like the sponges, too, most of the members of this branch are fixed, and in their general appearance suggest a plant rather than an animal. The name zoophytes, or plant-animals, which is often applied to these animals is based on this superficial resemblance. But many of the Cœlenterates lead an active free-swimming life. This is true of the jellyfishes which float or swim about on or near the surface of the ocean. Many of the zoophytes spend part of their life in an active free-swimming condition before settling down, becoming attached and thereafter remaining fixed. In localities near the seashore many animals belonging to this great group can be readily found and observed. The beautiful sea-anemones with their slowly-waving tentacles, the fine many-branched truly plant-like hydroids with their hosts of little buds, and the soft colorless masses of jelly, the jellyfishes, which are cast up on to the beaches by the waves are all animals belonging to the branch Cœlenterata.

General form and organization of body.—The general or typical plan of body-structure for the Cœlenterata, these animals which come next to the sponges in degree of complexity, can best be understood by imagining the typical cylindrical or vase-like body of the simple sponges to be modified in the following way: The middle one of the three layers of the body-wall not to be composed of scattered cells in a gelatinous matrix, but to be simply a thin non-cellular membrane; the body-wall not to be pierced by fine openings or pores, but connected with the outside only by the single large opening at the free end, and this opening to be surrounded by a circlet of arm-like processes or tentacles, which are continuations of the body-wall and similarly composed. Such a body-structure, which we saw well shown by Hydra, is the fundamental one for all polyps, sea-anemones, corals, and jellyfishes. The variety in shape of the body and the superficial modifications of this type-plan are many and striking, but after all the type-plan is recognizable throughout the whole of this great group of animals.

The two chief body-shapes represented in the branch are those of the polyps on the one hand, and the jellyfishes or medusæ on the other. The polyp-shape is that of a tube with a basal end blind or closed, attached to some firm object in the water and with the free end with an opening, the mouth-opening. At this mouth-end there is a circlet of movable, very contractile tentacles. The mouth may open directly into the interior of the body, which interior may be called the digestive cavity, or it may lead into a simple short tube produced by the invagination or bending in of the body-wall, which may be looked on as the simplest kind of œsophagus. This œsophageal tube opens into the body-cavity or digestive cavity. This cavity may be incompletely divided by longitudinal partitions which project from the sides into the cavity.

The jellyfish or medusoid body-form corresponds in general to an umbrella or bell. Around the edge of this umbrella are disposed numerous threads or tentacles (corresponding to the circlet of tentacles in the polyp). The mouth-opening is at the end of a longer or shorter projection which hangs down from the middle of the under side of the umbrella. The interior body-cavity or digestive cavity extends out into the umbrella-shaped part of the body, usually in the condition of canals radiating from the centre and a connecting canal running around the margin of the umbrella.

Structure.—Although the Cœlenterata show little indication of the complex composition of the body out of organs, as it exists among the higher animals, yet they do show an unmistakable advance on the simple, almost organless body of the sponges. This is chiefly shown by the differentiation among the cells which compose the body. In the polyps and jellyfishes some of the cells are specialized to be unmistakable muscle-cells, some to be nerve-cells and fibres, and so on. A very simple nervous system consisting of small groups of nerve-cells connected by nerve-fibres exists. Some very simple special sense-organs may occur. The digestive system, although in the simpler Cœlenterates consisting merely of the cylindrical body-cavity enclosed by the body-wall and opening by the single hole at the free end of the body, in some is rather complex and is composed of different parts. Those Cœlenterates which are not fixed but lead an active, free-swimming life, viz., the jellyfishes or medusæ, are the most highly organized.

The tentacles which surround the mouth-opening and serve to grasp food and carry it into the mouth, and the stinging or lasso threads with which these tentacles are provided are special organs possessed by most of these animals.

Skeleton.—Like the sponges, some of the Cœlenterata possess a hard skeleton. This skeleton is always composed of calcium carbonate and is called coral. Those polyps which form such a skeleton are called the corals. Coral will be described in connection with the account of the coral-polyps.