What families, either through their mass of individuals or their number of species, take precedence of the other Phanerogams?
How many species of one and the same family belong to any particular country?
What groups or families are characteristic of each zone?
Is the present classification of genera and species in all respects what could be desired?
These are questions that require to be considered, and to some of them we shall presently attempt replies.
Herbariums, though their classification is too frequently imperfect, may furnish us with data of great utility. The great herbarium of Benjamin Delessert was estimated, after his death, to contain 86,000 species,—a total not widely differing from that which Lindley, in 1835, estimated as the probable aggregate of the vegetable species of the world.
Great in importance are botanic gardens. Loudon, in his Hortus Britannicus (ed. 1832), places at 22,660 the number of Phanerogams cultivated in the gardens of the Bristol amateur botanists. With this number we must not confound the living species exhibited, in other counties, in gardens designed for the instruction of students, nor the grand total reared for a similar purpose at Kew. Kunth's enumeration, in 1861, of the plants at the Botanic Gardens at Berlin, one of the richest in Europe, amounted to upwards of 14,000 species, including 375 heaths. Among the Phanerogams were 1600 Synantheræ, 1150 Leguminosæ, 428 Labiatæ, 370 Umbelliferæ, 460 Orchidaceæ, 60 Palmaceæ, 600 Gramineæ and Cyperaceæ, &c. By comparing these data with the number of species described in the works of De Candolle, Walpers, Bentham, Lindley, Kunth, and others, we find that in the Berlin gardens are cultivated only one-seventh of the known species of the Synantheræ, one-eighth of the Leguminosæ, one-ninth of the Gramineæ, and about one-fiftieth of the smaller families, such as the Labiatæ and Umbelliferæ.
Now, if we admit that, on the one hand, the number of phanerogamous species cultivated in all the great gardens of Europe is about 30,000, and, on the other, that the cultivated Phanerogams form about one-eighth part of the species described in books and preserved in herbariums, we obtain a total of 24,000 species.
But the Cryptogams, or Agams, such as heaths, mosses, lichens, mushrooms, fungi, mould, and the like, of which our knowledge, as yet, is very imperfect, are probably much more numerous in species than the Phanerogams; for these vegetables, mostly microscopical, develop themselves wherever life can manifest itself—on the barren and denuded rocks, as well as in the air and in the depths of the ocean. If we suppose that they exceed only by 2000 the estimated number of Phanerogams, we shall obtain a total of just half-a-million!