This brief sketch of the cranial characters of the Krapina remains must be supplemented by a note on the teeth. Great numbers were found, and some of them are of enormous dimensions, surpassing those of the Mauer jaw. But some of the molar teeth are further distinguished in a very remarkable way, for the roots supporting the crown of the tooth are conjoined or fused: they are not distinct or divergent as is usual. The contrast thus provided by these anomalous teeth is well illustrated in the accompanying figure (8, Ko). Now such fusion of roots is not absolutely unknown at the present day; but the third molar or wisdom tooth is most frequently affected. The occurrence is extremely unusual in the other molar teeth of modern men. Yet among the Krapina teeth, such fusion is striking both in its degree and in its frequency. So marked a characteristic has attracted much attention. Professor Kramberger holds the view that it constituted a feature of adaptation peculiar to the Palaeolithic men of Krapina. In opposition to this, Professor Adloff holds that the character is so definite and marked as to enter into the category of distinctive and specific conformations. The discussion of these views was carried on somewhat warmly, but yet to some extent fruitlessly so long as the only known examples were those from Krapina. Dr Laloy supported Professor Kramberger, and on the other side may be ranged the support of Professor Walkhoff. But a recent discovery has very substantially fortified the view adopted by Professor Adloff and his supporters. For in a cave near S. Brélade's Bay in Jersey, the explorations of Messrs Nicolle, Sinel and Marett (1910-1911) have brought to light Palaeolithic human teeth of very similar form. They are said indeed by Dr Keith to be precisely comparable to those from Krapina. The conjoined roots of such teeth should be regarded therefore as more than a peculiarity of the Palaeolithic men of Croatia, and rather as a very definite means of assigning to a particular Palaeolithic epoch any other instances of a similar nature. Space will not admit of more than a simple record of two other features of the Krapina teeth. They are (a) the curvature of the canine teeth and (b) the remarkable size and extent of the ‘pulp-cavity’ (cf. [Fig. 8], Ko) of the molar teeth. In entering upon so protracted a discussion of this part of the evidence, the excuse is proffered that, as may be noted in the instances at Trinil and Taubach, teeth are remarkably well-fitted for preservation in the fossil state, since they may be preserved in circumstances leading to the complete destruction of other parts of the skeleton.
The limb bones of the Krapina skeletons are chiefly remarkable for the variety they present. Some are short and stout, of almost pygmy proportions: others are long and slender, inappropriate in these respects to the massive skull fragments which predominate. The distinction of two human types upon evidence furnished by the limb bones has already been mentioned.
S. Brélade's Bay, Jersey.
A cave in this locality has been explored during the last two years (1910, 1911). Human remains are represented by the teeth already mentioned on account of their resemblance to those found at Krapina. The resemblance depends primarily upon the curious fusion of the roots in the molar teeth. Moreover, the circumference of the combined and thickened roots is so great as to confer a most remarkable ‘columnar’ appearance on the affected teeth (cf. [fig. 8], K.o.). The teeth from Krapina and Jersey while thus associated must be contrasted with some specimens which they resemble in other respects. The corresponding teeth in the Mauer jaw have been described as similar to those from Krapina, but I cannot confirm this from Dr Schoetensack's illustrations, of which fig. 8 (H) is a fair representation. The teeth of the Forbes Quarry and Le Moustier specimens do not conform to the precise requirements of the test. The Spy teeth are said to have three distinct roots save in two cases, where the numbers are four and two respectively. The test of combined molar roots therefore provides a means of subdividing a group of examples otherwise similar, rather than a mark of recognition applicable to all alike.
The S. Brélade teeth also resemble those from Krapina in the proportions of their crowns and the unusually large size of the pulp-cavity. The latter character may prove more important than the fusion of the roots. But the evidence of their surroundings assigns the teeth from Jersey to an epoch less ancient than that of the Krapina men.
La Chapelle-aux-Saints (Corrèze).
The human skeleton from La Chapelle-aux-Saints holds a very distinguished position among its congeners. In the first place, the discovery was not haphazard, but made by two very competent observers during their excavations. Again, the remains comprise not only the nearly intact brain-case, but much of the facial part of the skull, together with the lower jaw and many bones of the trunk and limbs. The individual was a male of mature age, but not senile (Manouvrier). For these reasons, the value of this skeleton in evidence is singularly great.