To sum up, it will be found that the evidence of the Elephant is to the effect that these alluvial deposits are of early Pleistocene age. It leads to the expectation that the fauna in general will have a ‘southern,’ as contrasted with an ‘arctic’ aspect. From the study of the Rhinoceros it appears that the Mauer Sands are probably the most ancient in order of time, that the strata of Taubach are the latest of the three and that Elephas antiquus will occur there (as indeed it does).

The other animals mentioned clinch the evidence for the Pliocene resemblance, and (at latest) the early Pleistocene antiquity of the Mauer Sands and the high-level terrace gravels. Within the limits thus indicated, the deposit of Mauer is again shewn to be the oldest, followed by the terrace-gravels, while Taubach is the latest and youngest of the three. All the characteristic animals are now entirely extinct.

For the reasons stated above, the fossil Javanese mammals of Trinil have not been discussed. It will suffice to note that on the whole they indicate a still earlier period than those of the European deposits in question.

(ii) The animals associated with the cave-men now call for consideration. The great outstanding feature is the constancy with which the Reindeer is found. This leads to a presumption that the climate was at least temperate rather than ‘southern.’ Beyond this, it will be noted that in general the cave-fauna is more familiar in aspect, the Reindeer having survived up to the present day, though not in the same area. Again, save in one locality, not a single animal out of those discussed in connection with the alluvial deposits appears here. The exception is the Krapina rock-shelter. The surviving animal is Rhinoceros merckii, described above as one of the later arrivals in the epochs represented by the alluvial deposits. Krapina does not provide the Reindeer, and in this respect is contrasted again with the remaining localities. Yet the presence of the Marmot at Krapina may be nearly as significant as that of the Reindeer would be.

Another cave, viz. the Grotte des Enfants, may also need reconsideration. For instance, the Rhinoceros merckii was found in the deepest strata of this cave: but I do not consider that adequate evidence is given of its contemporaneity with the two human skeletons here considered. But the Reindeer is found in the same cave, as indicated in the table.

With the exception of Krapina therefore, the conditions are remarkably uniform. This conclusion is confirmed by the evidence from many caves not described in detail here because of the lack of human bones therein or the imperfection of such as were found. Such caves have yielded abundant evidence in regard to the ‘associated fauna.’ A few of the more important results of the investigation of the mammals may be given. Thus the distribution of the Reindeer is so constant that except in regard to its abundance or rarity when compared with the remains of the horse in the same cave, it is of little or no use as a discriminating agency. The Mammoth (E. primigenius) was contemporaneous with the Reindeer, but was plentiful while the Reindeer was still rare. A similar remark applies to the Hairy Rhinoceros (R. tichorhinus), and also to the Cave-Bear. The Cervidae (other than the Reindeer), the Equidae, the Suidae (Swine) and the smaller Rodentia (especially Voles) are under investigation, but the results are not applicable to the finer distinctions envisaged here.

To sum up the outcome of this criticism; it appears that of the cave-finds, Krapina stands out in contrast with the remainder, in the sense that its fauna is more ancient, and is indicative of a southern rather than a temperate environment. The latitude of Krapina has been invoked by way of explaining this difference, upon the supposition that the Rhinoceros merckii survived longer in the south. Yet Krapina does not differ in respect of latitude from the caves of Le Moustier and La Chapelle, while it is rather to the north of the Mentone caves. Lastly, some weight must be attached to the alleged discovery at Pont Newydd in Wales, of Mousterian implements with remains of R. merckii.

The fauna of the other caves suggests temperate, if not sub-arctic conditions of climate. In all cases, the cave-finds are assignable to a period later in time than that in which the fluviatile deposits (previously discussed) were formed. The cave-men thus come within the later subdivisions of the Pleistocene period.

(c) The fifth column of the table gives the types of stone implements found in association with the respective remains. As is well known, and as was stated in the introductory sentences of this book, stone artefacts constitute the second great class of evidence on the subject of human antiquity. As such they might appropriately have been accorded a separate chapter or even a volume. Here a brief sketch only of their significance in evidence will be attempted. The value of stone implements in deciding upon the age of deposits (whether in caves or elsewhere) depends upon the intimacy of the relation existing between various forms of implement and strata of different age. How close that intimacy really is, has been debated often and at great length. Opinions are still at variance in regard to details, but as to certain main points, no doubt remains. Yet the study is one in which even greater specialisation is needed than in respect of comparative osteology. The descriptions following these preliminary remarks are based upon as extensive an examination as possible, both of the literature, and of the materials.

To discuss the validity of the claims made in favour of or against the recognition of certain individual types will be impossible, save in the very briefest form. The better-known varieties have received names corresponding to the localities where they were first discovered, or where by reason of their abundance they led to the recognition of their special value as a means of classification. These designations will be employed without further definition or explanation, save in a few instances.