In what sense the observations of the two men were complemental will be better understood when we remember that there are two aspects in which living organisms should always be considered in biological studies; first, the structure, and, then, the use that the structures subserve. One view is essential to the other, and no investigation of animals and plants is complete in which the two ideas are not involved. Just as a knowledge of the construction of a machine is necessary to understand its action, so the anatomical analysis of an organ must precede a knowledge of its office. The term "physiological anatomy of an organ," so commonly used in text-books on physiology, illustrates the point. We can not appreciate the work of such an organ as the liver without a knowledge of the arrangement of its working units. The work of the anatomist concerns the statics of the body, that of the physiologist the dynamics; properly combined, they give a complete picture of the living organism.

It is to be remembered that the observations of Vesalius were not confined exclusively to structure; he made some experiments and some comments on the use of parts of the body, but his work was mainly structural, while that which distinguishes Harvey's research is inductions founded on experimental observation of the action of living tissues.

The service of Vesalius and Harvey in opening the path to biological advance is very conspicuous, but they were not the only pioneers; their work was a part of the general revival of science in which Galileo, Descartes, and others had their part. While the birth of the experimental method was not due to the exertions of Harvey alone, nevertheless it should stand to his credit that he established that method in biological lines. Aristotle and Galen both had employed experiments in their researches, and Harvey's step was in the nature of a revival of the method of the old Greeks.

Harvey's Education.—Harvey was fitted both by native talent and by his training for the part which he played in the intellectual awakening. He was born at Folkestone, on the south coast of England, in 1578, the son of a prosperous yeoman. The Harvey family was well esteemed, and the father of William was at one time the mayor of Folkestone. Young Harvey, after five years in the King's school at Canterbury, went to Cambridge, and in 1593, at the age of sixteen, entered Caius College. He had already shown a fondness for observations upon the organization of animals, but it is unlikely that he was able to cultivate this at the university. There his studies consisted mainly of Latin and Greek, with some training in debate and elementary instruction in the science of physics.

At Padua.—In 1597, at the age of nineteen, he was graduated with the Arts degree, and the following year he turned his steps toward Italy in search of the best medical instruction that could be found at that time in all the world. He selected the great university of Padua as his place of sojourn, being attracted thither by the fame of some of its medical teachers. He was particularly fortunate in receiving his instruction in anatomy and physiology from Fabricius, one of the most learned and highly honored teachers in Italy. The fame of this master of medicine, who, from his birthplace, is usually given the full name of Fabricius ab Aquapendente, had spread to the intellectual centers of the world, where his work as anatomist and surgeon was especially recognized. A fast friendship sprang up between the young medical student and this ripe anatomist, the influence of which must have been very great in shaping the future work of Harvey.

Fabricius was already sixty-one years of age, and when Harvey came to Padua was perfecting his knowledge upon the valves of the veins. The young student was taken fully into his confidence, and here was laid that first familiarity with the circulatory system, the knowledge of which Harvey was destined so much to advance and amplify. But it was the stimulus of his master's friendship, rather than what he taught about the circulation, that was of assistance to Harvey. For the views of Fabricius in reference to the circulation were those of Galen; and his conception of the use of the valves of the veins was entirely wrong. A portrait of this great teacher of Harvey is shown in Fig. 9.

At Padua young Harvey attracted notice as a student of originality and force, and seems to have been a favorite with the student body as well as with his teachers. His position in the university may be inferred from the fact that he belonged to one of the aristocratic-student organizations, and, further, that he was designated a "councilor" for England. The practice of having student councilors was then in vogue in Padua; the students comprising the council met for deliberations, and very largely managed the university by their votes upon instructors and university measures.

It is a favorable comment upon the professional education of his time that, after graduating at the University of Cambridge, he studied four or more years (Willis says five years) in scientific and medical lines to reach the degree of Doctor of Physic.

On leaving Padua, in 1602, he returned to England and took the examinations for the degree of M.D. from Cambridge, inasmuch as the medical degree from an English university advanced his prospects of receiving a position at home. He opened practice, was married in 1604, and the same year began to give public lectures on anatomy.