The most significant advances in reference to the classification of animals was to come as a result of the acceptance of the doctrine of organic evolution, subsequent to 1859. Then the relationships between animals were made to depend upon community of descent, and a distinction was drawn between superficial or apparent relationships and those deep-seated characteristics that depend upon close genetic affinities.

Alterations by Von Siebold and Leuckart.—But, in the mean time, naturalists were not long in discovering that the primary divisions established by Cuvier were not well balanced, and, indeed, that they were not natural divisions of the animal kingdom. The group Radiata was the least sharply defined, since Cuvier had included in it not only those animals which exhibit a radial arrangement of parts, but also unicellular organisms that were asymmetrical, and some of the worms that showed bilateral symmetry. Accordingly, Karl Th. von Siebold, in 1845, separated these animals and redistributed them. For the simplest unicellular animals he adopted the name Protozoa, which they still retain, and the truly radiated forms, as starfish, sea-urchins, hydroid polyps, coral animals, etc., were united in the group Zoöphyta. Von Siebold also changed Cuvier's branch, Articulata, separating those forms as crustacea, insects, spiders, and myriopods, which have jointed appendages, into a natural group called Arthropoda, and uniting the segmented worms with those worms that Cuvier has included in the radiate group, into another branch called Vermes. This separation of the four original branches of Cuvier was a movement in the right direction, and was destined to be carried still farther.

Fig. 35.—Karl Th. von Siebold, 1804-1885.

Von Siebold (Fig. 35) was an important man in the progress of zoölogy, especially in reference to the comparative anatomy of the invertebrates.

Leuckart (Fig. 36), whose fame as a lecturer and teacher attracted many young men to the University of Leipsic, is another conspicuous personality in zoölogical progress.

This distinguished zoölogist, following the lead of Von Siebold, made further modifications. He split Von Siebold's group of Zoöphytes into two distinct kinds of radiated animals; the star-fishes, sea-urchins, sea-cucumbers, etc., having a spiny skin, he designated Echinoderma; the jelly-fishes, polyps, coral animals, etc., not possessing a true body cavity, were also united into a natural group, for which he proposed the name Cœlenterata.