A diagram of the cell as now understood (Fig. 80) will be helpful in showing how much the conception of the cell has changed since the time of Schleiden and Schwann.
Definition.—The definition of Verworn, made in 1895, may be combined with this diagram: A cell is "a body consisting essentially of protoplasm in its general form, including the unmodified cytoplasm, and the specialized nucleus and centrosome; while as unessential accompaniments may be enumerated: (1) the cell membrane, (2) starch grains, (3) pigment granules, (4) oil globules, and (5) chlorophyll granules." No definition can include all variations, but the one quoted is excellent in directing attention to the essentials—to protoplasm in its general form, and the modified protoplasmic parts as distinguished from the unessential accompaniments, as cell membrane and cell contents.
The definition of Verworn was reached by a series of steps representing the historical advance of knowledge regarding the cell. Schleiden and Schwann looked upon the cell as a hollow chamber having a cell-wall which had been formed around the nucleus; it was a great step when Schultze defined the cell in terms of living substance as "a globule of protoplasm surrounding a nucleus," and it is a still deeper level of analysis which gives us a discriminating definition like that of Verworn.
When we are brought to realize that, in large part, the questions that engage the mind of the biologist have their basis in the study of cells, we are ready to appreciate the force of the statement that the establishment of the cell-theory was one of the great events of the nineteenth century, and, further, that it stands second to no theory, with the single exception of that of organic evolution, in advancing biological science.
PROTOPLASM, THE PHYSICAL BASIS OF LIFE
The recognition of the rôle that protoplasm plays in the living world was so far-reaching in its results that we take up for separate consideration the history of its discovery. Although it is not yet fifty years since Max Schultze established the protoplasm doctrine, it has already had the greatest influence upon the progress of biology. To the consideration of protoplasm in the previous chapter should be added an account of the conditions of its discovery, and of the personality and views of the men whose privilege it was to bring the protoplasm idea to its logical conclusion. Before doing so, however, we shall look at the nature of protoplasm itself.