With piebalds the same result is asserted, save that certain melanic forms appeared. Finally von Fischer repeats his laws already reached, giving them now in this form: that if the offspring of a cross show only the colour of the father, then the parents are varieties of one species; but if the colour of the offspring be intermediate or different from that of the father, then the parents belong to distinct species.
The reader may have already gathered that we have here that bane of the advocate—the witness who proves too much. But why does Professor Weldon confine von Fischer to the few modest words recited above? That author has—so far as colour is concerned—a complete law of heredity supported by copious “observations.” Why go further?
Professor Weldon “brings forth these strong reasons” of the rats and mice with the introductory sentence:
“Examples might easily be multiplied, but as before, I have chosen rather to cite a few cases which rest on excellent authority, than to quote examples which may be doubted. I would only add one case among animals, in which the evidence concerning the inheritance of colour is affected by the ancestry of the varieties used.”
So once again Professor Weldon suggests that his laws of ancestry will explain even the discrepancies between von Fischer on the one hand and Crampe and von Guaita on the other but he does not tell us how he proposes to apply them.
In the cross between the albino and the grey von Fischer tells us that both colours appear in the offspring, but always, without exception or variation, that of the father only, in 5847 individuals.
Surely, the law of ancestry, if he had a moment’s confidence in it, might rather have warned Professor Weldon that von Fischer’s results were wrong somewhere, of which there cannot be any serious doubt. The precise source of error is not easy to specify, but probably carelessness and strong preconception of the expected result were largely responsible, though von Fischer says he did all the recording most carefully himself.
Such then is the evidence resting “on excellent authority”: may we some day be privileged to see the “examples which may be doubted”?
The case of mice, invoked by Professor Weldon, has also been referred to in our Report. Its extraordinary value as illustrating Mendel’s principles and the beautiful way in which that case may lead on to extensions of those principles are also there set forth (see the present Introduction, p. [25]). Most if not all of such “conflicting” evidence can be reconciled by the steady application of the Mendelian principle that the progeny will be constant when—and only when[138]—similar gametes meet in fertilisation, apart from any question of the characters of the parent which produces those gametes.