A history of land mammals in the western hemisphere

by William Berryman Scott

In working out the geographical conditions for any particular epoch of the earth’s history, it is possible to go much farther than merely gaining an approximate estimate of the distribution of land and sea; many other important facts may be gathered from a minute examination of the rocks in combination with a genetic study of topographical forms. By this physiographical method, as it is called, the history of several of the great mountain-ranges has been elaborated in great detail. It is quite practicable to give a geological date for the initial upheaval and to determine whether one or many such series of movements have been involved in bringing about the present state of things. Similarly, the history of plains and plateaus, hills and valleys, lake and river systems, may be ascertained, and for the earth’s later ages, at least, a great deal may be learned regarding the successive forms of the land-surfaces in the various continents. It would be very desirable to explain the methods by which these results are reached, but this could hardly be done without writing a treatise on physiography, for which there is no room in this chapter. We must be permitted to make use of the results of that science without being called upon to prove their accuracy.

No factor has a more profound effect in determining the character and distribution of living things than climate, of which the most important elements, for our purpose, are temperature and moisture. One of the most surprising results of geological study is the clear proof that almost all parts of the earth have been subjected to great vicissitudes of climate, and a brief account of the evidence which has led to this unlooked for result will not be out of place here.

The evidence of climatic changes is of two principal kinds, (1) that derived from a study of the rocks themselves, and (2) that given by the fossils of the various epochs. So far as the rocks laid down in the sea are concerned, little has yet been ascertained regarding the climatic conditions of their formation, but the strata which were deposited on the land, or in some body of water other than the sea, often give the most positive and significant information concerning the circumstances of climate which prevailed at the time of their formation. Certain deposits, such as gypsum and rock-salt, are accumulated only in salt lakes, which, in turn, are demonstrative proof of an arid climate. A salt lake could not exist in a region of normal rainfall and, from the geographical distribution of such salt-lake deposits, it may be shown that arid conditions have prevailed in each of the continents and, not only once, but many times. As a rule, such aridity of climate was relatively local in extent, but sometimes it covered vast areas. For example, in the Permian, the last of the Palæozoic, and the Triassic, the first of the Mesozoic periods (see Table, [p. 15]) nearly all the land-areas of the northern hemisphere were affected, either simultaneously or in rapid succession.

Until a comparatively short time ago, it was very generally believed that the Glacial or Pleistocene epoch, which was so remarkable and conspicuous a feature of the Quaternary period, was an isolated phenomenon, unique in the entire history of the earth. Now, however, it has been conclusively shown that such epochs of cold have been recurrent and that no less than five of these have left unmistakable records in as many widely separated periods of time.

When the hypothesis of a great “Ice Age” in the Pleistocene was first propounded by the elder Agassiz, it was naturally received with general incredulity, but the gradual accumulation of proofs has resulted in such an overwhelming weight of testimony, that the glacial hypothesis is now accepted as one of the commonplaces of Geology. The proofs consist chiefly in the characteristic glacial accumulations, moraines and drift-sheets, which cover such enormous areas in Europe and North America and, on a much smaller scale, in Patagonia, and in the equally characteristic marks of glacial wear left upon the rocks over which the ice-sheets moved. Many years later it was proved that the Permian period had been a time of gigantic glaciation, chiefly in the southern hemisphere, when vast ice-caps moved slowly over parts of South America, South Africa, Australia and even of India. The evidence is of precisely the same nature as in the case of the Pleistocene glaciation. In not less than three more ancient periods, the Devonian, Cambrian, and Algonkian, proofs of glacial action have been obtained.

While the rocks themselves thus afford valuable testimony as to the climatic conditions which prevailed at the time and place of their formation, this testimony is fragmentary, missing for very long periods, and must be supplemented from the information presented by the fossils. As in all matters where fossils are involved, the evidence must be cautiously used, for hasty inferences have often led to contradictory and absurd conclusions. When properly employed, the fossils give a more continuous and complete history of climatic changes than can, in the present state of knowledge, be drawn from a study of the rocks alone. For this purpose plants are particularly useful, because the great groups of the vegetable kingdom are more definitely restricted in their range by the conditions of temperature and moisture than are most of the correspondingly large groups of animals. Not that fossil animals are of no service in this connection; quite the contrary is true, but the evidence from them must be treated more carefully and critically. To illustrate the use of fossils as recording climatic changes in the past, one or two examples may be given.

In the Cretaceous period a mild and genial climate prevailed over all that portion of the earth whose history we know, and was, no doubt, equally the case in the areas whose geology remains to be determined. The same conditions extended far into the Arctic regions, and abundant remains of a warm-temperate vegetation have been found in Greenland, Alaska and other Arctic lands. Where now only scanty and minute dwarf willows and birches can exist, was then a luxuriant forest growth comprising almost all of the familiar trees of our own latitudes, a most decisive proof that in the Cretaceous the climate of the Arctic regions must have been much warmer than at present and that there can have been no great accumulation of ice in the Polar seas. Conditions of similar mildness obtained through the earlier part of the Tertiary. In the Eocene epoch large palm-trees were growing in Wyoming and Idaho, while great crocodiles and other warm-country reptiles abounded in the waters of the same region.

It is of particular interest to inquire how far the fossils of Glacial times confirm the inferences as to a great climatic change which are derived from a study of the rocks, for this may be taken as a test-case. Any marked discrepancy between the two would necessarily cast grave doubt upon the value of the testimony of fossils as to climatic conditions. The problem is one of great complexity, for the Pleistocene was not one long epoch of unbroken cold, but was made up of Glacial and Interglacial ages, alternations of colder and milder conditions, and some, at least, of the Interglacial ages had a climate warmer than that of modern times. Such great changes of temperature led to repeated migrations of the mammals, which were driven southward before the advancing ice-sheets and returned again when the glaciers withdrew under the influence of ameliorating climates. Any adequate discussion of these complex conditions is quite out of the question in this place and the facts must be stated in simplified form, as dealing only with the times of lowered temperature and encroaching glaciers.

The plants largely fail us here, for little is known of Glacial vegetation, but, on the other hand, a great abundance of the fossil remains of animal life of that date has been collected, and its testimony is quite in harmony with that afforded by the ice-markings and the ice-made deposits. Arctic shells in the marine deposits of England, the valley of the Ottawa River and of Lake Champlain, Walruses on the coast of New Jersey, Reindeer in the south of France, and Caribou in southern New England, Musk-oxen in Kentucky and Arkansas, are only a few examples of the copious evidence that the climate of the regions named in Glacial times was far colder than it is to-day.

I have thus endeavored to sketch, necessarily in very meagre outlines, the nature of the methods employed to reconstruct the past history of the various continents and the character of the evidence upon which we must depend. Should the reader be unconvinced and remain sceptical as to the possibility of any such reconstruction, he must be referred to the numerous manuals of Geology, in which these methods are set forth with a fulness which cannot be imitated within the limits of a single chapter. The methods are sound, consisting as they do merely in the application of “systematized common sense” (in Huxley’s phrase) to observed facts, but by no means all applications of them are to be trusted. Not to mention ill-considered and uncritical work, or inverted pyramids of hypothesis balanced upon a tiny point of fact, it should be borne in mind that such a complicated and difficult problem as the reconstruction of past conditions can be solved only by successive approximations to the truth, each one partial and incomplete, but less so than the one which preceded it.