A history of land mammals in the western hemisphere

by William Berryman Scott

In the interior, the Pliocene continental formations and faunas followed so gradually upon those of the Miocene, that there is great doubt as to where the line between them should be drawn. These interior formations are mostly of small extent and are very widely scattered, and much remains to be learned regarding the mammals of the epoch. In northern Kansas are the Republican River beds, which are so doubtfully Pliocene, that they may almost equally well be called uppermost Miocene. Other lower Pliocene stages, representing various divisions of time, are the Alachua of northern Florida, the Snake Creek of western Nebraska, the Thousand Creek and Virgin Valley of northwestern Nevada and the Rattlesnake of Oregon. Probably middle Pliocene is the Blanco of northwestern Texas, a valley cut in the middle and lower Miocene rocks and filled in with Pliocene deposits. Possibly upper Pliocene, or, it may be, lowest Pleistocene, are the Peace Creek of southwestern Florida and the so-called “Loup River” (not Loup Fork) of western Nebraska.

The volcanic activity of the Rocky Mountain and Pacific coast regions, which was so remarkable in the Miocene, continued into and perhaps through the Pliocene. The great outflow of light-coloured lava which built up the central plateau of the Yellowstone Park is referred to the Pliocene, and some of the enormous fissure-eruptions which formed the vast Columbia River fields of black basaltic lava were probably Pliocene, as some were demonstrably Miocene. Both of these epochs were remarkable for volcanic activity in the western part of the continent.

The Pliocene climate, as may be inferred from the plants and marine shells, was colder than that of the Miocene, and refrigeration was progressive, as is shown by the proportion of Arctic shells in the Pliocene beds of the east coast of England, rising from 5 per cent in the oldest to more than 60 per cent in the latest beds. In the Arctic regions the cold must have been severe, at least during the latter half of the epoch, for in the succeeding Pleistocene we find an Arctic fauna already fully adapted to the extreme severity of present day polar conditions and time was necessary for such an adaptation. In the western interior the climate was not only colder, but also drier than it had been in the Miocene, the desiccation which had begun in the latter epoch becoming progressively more and more marked.

South America.—The Greater Antilles were larger than at present and Cuba was much extended, especially to the southeastward, and was probably connected with the mainland, not as one would naturally expect, with Yucatan, but with Central America; this island, it is most likely, was cut off from Hayti. The Isthmian region was considerably broader than it is now and afforded a more convenient highway of intermigration. Costa Rica was invaded by a Pliocene gulf, but it is not yet clear whether this persisted for the whole or only a part of the epoch. In the Argentine province of Entrerios is a formation, the Paraná, which is most probably Pliocene, though it may be upper Miocene. This formation is largely marine and shows that the present Rio de la Plata was then a gulf from the Atlantic. A few northern hemisphere mammals in the Paraná beds show that the migration had advanced far into South America. A large part of Patagonia was again submerged beneath the sea, which extended to the Andes in places, but just how general the submergence was, it is impossible to say, for the Cape Fairweather formation has been largely carried away by erosion and only fragments of it remain. Along the foothills of the Andes these beds are upturned and raised several thousand feet above the sea-level, a proof that the final upheaval of the southern mountains took place at some time later than the early Pliocene. Continental formations of Pliocene date are largely developed in Argentina; the Araucanian stage is in two substages, one in the province of Catamarca, where the beds are much indurated and were involved in the Andean uplift, the other, of unconsolidated materials, is at Monte Hermoso near Bahia Blanca on the Atlantic coast. The very small proportion of northern animals in the Araucanian beds is surprising, but not more so than the almost complete absence of South American types in the upper Miocene and lower Pliocene of the United States. Intermigration between the two Americas would seem to have been a much slower and more difficult process than between North America and the Old World, and the reason for the difference is probably the greater climatic barriers involved in a migration along the lines of longitude. Upper Pliocene is found in the Tarija Valley of Bolivia and probably also in Ecuador, in both of which areas the proportion of northern animals was very greatly increased.

II. Quaternary Period

The Quaternary period was a time of remarkable geographical and climatic changes, which had the profoundest and most far-reaching effects, partly by migration and partly by extinction, upon the distribution of animals and plants, effects which are naturally more obvious than those of earlier geological events, just because they were the latest. It is customary to divide the period into two epochs, (1) the Pleistocene or Glacial, and (2) the Recent, which continues to the present day.

1. Pleistocene Epoch

When Louis Agassiz first suggested (1840) the idea of a time, comparatively recent in the geological sense, when northern and central Europe was buried under immense sheets of slowly moving ice, like the “ice-cap” of modern Greenland, the conception was received with incredulity. Nearly thirty years passed before this startling theory gained the general acceptance of geologists, but now it is one of the commonplaces of the science, for no other hypothesis so well explains the complicated phenomena of Pleistocene geology. One great obstacle to the acceptance of the glacial theory was the supposed fact that the Pleistocene glaciation was something quite unique in the history of the earth, a violent aberration in the development of climates. Now, however, we have every reason to believe that at least three other and very ancient periods had witnessed similar climatic changes and that “ice-ages” were recurrent phenomena. This is not the place to discuss or even to summarize the evidence which has convinced nearly all geologists of the reality of Pleistocene glacial conditions on a vast scale in Asia, Europe and, above all, in North America. The reader who may wish to examine this evidence will find an admirable presentation of it in Vol. III of the “Geology” of Professors Chamberlin and Salisbury.

North America.—There has long been a difference of opinion among students of the Pleistocene as to whether the glaciation was single, or several times renewed. That there were many advances and retreats of the ice, is not denied; the question is, whether there were truly interglacial stages, when the ice altogether disappeared from the continent and the climate was greatly ameliorated. The present tendency among American and European geologists is decidedly in favour of accepting several distinct glacial stages (Chamberlin and Salisbury admit six of these) separated by interglacial stages, and for this there are very strong reasons. While it is out of the question to present the evidence for this conclusion here, one or two significant facts may be noted. On the north shore of Lake Ontario, near Toronto, are certain water-made deposits, which rest upon one sheet of glacial drift and are overlaid by another. The fossils of the aqueous sediments are in two series, upper and lower, of which the older and lower contains plants and insects indicative of a climate considerably warmer than that of the same region to-day and corresponding to the temperature of modern Virginia. In the upper and newer beds the fossils show the return of cold conditions, much like those of southern Labrador, and this was followed by the reëstablishment of the ice, as recorded in the upper sheet of drift. Even far to the north, on the Hudson’s Bay slope, an interglacial forest is embedded between two glacial drift-sheets. In Iowa and South Dakota numerous mammals of temperate character occur in interglacial beds.

At the time of their greatest extension, the glaciers covered North America down to latitude 40° N., though the great terminal moraine, which marks the ice-front and has been traced across the continent from Nantucket to British Columbia, describes a very sinuous line. The ice was not a homogeneous sheet, moving southward as a whole, but flowed in all directions away from several, probably four, centres of accumulation and dispersal. At the same time, the western mountain ranges had a far greater snow-supply than at present, and great glaciers flowed down all the valleys of the Rocky Mountains as far south as New Mexico and in the Sierras to southern California, while the Wasatch, Uinta and Cascade ranges and those of British Columbia and Alaska were heavily glaciated, but, strange to say, the lowlands of Alaska were free from ice. During the periods of greatest cold the rain-belt was displaced far to the south of its normal position, bringing a heavy precipitation to regions which are now extremely arid. In the Great Basin were formed two very large lakes; on the east side, rising high upon the flanks of the Wasatch Mountains, was Lake Bonneville, the shrunken and pygmy remnant of which is the Salt Lake of Utah, and on the west side, in Nevada, was Lake Lahontan. Lake Bonneville, which was nearly two-thirds the size of Lake Superior, discharged northward into the Snake River, a tributary of the Columbia, but Lahontan had no outlet. Each of these lakes had two periods of expansion, with a time of complete desiccation between them.