A HISTORY OF LAND MAMMALS IN
THE WESTERN HEMISPHERE
CHAPTER I
METHODS OF INVESTIGATION—GEOLOGICAL
The term Mammal has no exact equivalent in the true vernacular of any modern language, the word itself, like its equivalents, the French Mammifère and the German Säugethier, being entirely artificial. As a name for the class Linnæus adopted the term Mammalia, which he formed from the Latin mamma (i.e. teat) to designate those animals which suckle their young; hence the abbreviated form Mammal, which has been naturalized as an English word. “Beast,” as employed in the Bible, and “Quadruped” are not quite the same as mammal, for they do not include the marine forms, such as whales, dolphins, seals, walruses, or the flying bats, and they are habitually used in contradistinction to Man, though Man and all the forms mentioned are unquestionably mammals.
In attempting to frame a definition of the term Mammal, it is impossible to avoid technicalities altogether, for it is the complete unity of plan and structure which justifies the inclusion of all the many forms that differ so widely in habits and appearance. Mammals are air-breathing vertebrates, which are warm-blooded and have a 4-chambered heart; the body cavity is divided into pleural and abdominal chambers by a diaphragm; except in the lowest division of the class, the young are brought forth alive and are always suckled, the milk glands being universal throughout the class. In the great majority of mammals the body is clothed with hair; a character found in no other animals. In a few mammals the skin is naked, and in still fewer there is a partial covering of scales. The list of characters common to all mammals, which distinguish them from other animals, might be indefinitely extended, for it includes all the organs and tissues of the body, the skeletal, muscular, digestive, nervous, circulatory, and reproductive systems, but the two or three more obvious or significant features above selected will suffice for the purposes of definition.
While the structural plan is the same throughout the entire class, there is among mammals a wonderful variety of form, size, appearance, and adaptation to special habits. It is as though a musician had taken a single theme and developed it into endless variations, preserving an unmistakable unity through all the changes. Most mammals are terrestrial, living, that is to say, not only on the land, but on the ground, and are herbivorous in habit, subsisting chiefly or exclusively upon vegetable substances, but there are many departures from this mode of life. It should be explained, however, that the term terrestrial is frequently used in a more comprehensive sense for all land mammals, as distinguished from those that are aquatic or marine. Monkeys, Squirrels, Sloths and Opossums are examples of the numerous arboreal mammals, whose structure is modified to fit them for living and sleeping in the trees, and in some, such as the Sloths, the modification is carried so far that the creature is almost helpless on the ground. Another mode of existence is the burrowing or fossorial, the animal living partly or mostly, or even entirely underground, a typical instance of which is the Mole. The Beaver, Muskrat and Otter, to mention only a few forms, are aquatic and spend most of their life in fresh waters, though perfectly able to move about on the land. Marine mammals, such as the Seals and Whales, have a greatly modified structure which adapts them to life in the sea.
Within the limits of each of these categories we may note that there are many degrees of specialization or adaptation to particular modes of life. Thus, for example, among the marine mammals, the Whales and their allies, Porpoises, etc., are so completely adapted to a life in the seas that they cannot come upon the land, and even stranding is fatal to them, while the Seals frequently land and move about upon the shore. It should further be observed that mammals of the most diverse groups are adapted to similar modes of existence. Thus in one natural group or order of related forms, occur terrestrial, burrowing, arboreal and aquatic members, and the converse statement is of course equally true, that animals of similar life-habits are not necessarily related to one another, and very frequently, in fact, are not so related. Among the typically marine mammals, for example, there are at least three and probably four distinct series, which have independently become adapted to life in the sea.
Before attempting to set forth an outline of what has been learned regarding the history of mammalian life in the western hemisphere, it is essential to give the reader some conception of the manner in which that knowledge has been obtained. Without such an understanding of the methods employed in the investigation the reader can only blindly accept or as blindly reject what purports to be the logical inference from well-established evidence. How is that evidence to be discovered? and how may trustworthy conclusions be derived from it?
The first and most obvious step is to gather all possible information concerning the mammals of the present day, their structure (comparative anatomy), functions (physiology), and their geographical arrangement. This latter domain, of the geographical distribution of mammals, is one of peculiar significance. Not only do the animals of North America differ radically from those of Central and South America, but within the limits of each continent are more or less well-defined areas, the animals of which differ in a subordinate degree from those of other areas. The study of the modern world, however, would not of itself carry us very far toward the goal of our inquiries, which is an explanation, not merely a statement, of the facts. The present order of things is the outcome of an illimitably long sequence of events and can be understood only in proportion to our knowledge of the past. In other words, it is necessary to treat the problems involved in our inquiry historically; to trace the evolution of the different mammalian groups from their simpler beginnings to the more complex and highly specialized modern forms; to determine, so far as that may be done, the place of origin of each group and to follow out their migrations from continent to continent.
While we shall deal chiefly, almost exclusively, with the mammals of the New World, something must be said regarding those of other continents, for, as will be shown in the sequel, both North and South America have, at one time or another, been connected with various land-masses of the eastern hemisphere. By means of those land-connections, there has been an interchange of mammals between the different continents, and each great land-area of the recent world contains a more or less heterogeneous assemblage of forms of very diverse places of origin. Indeed, migration from one region to another has played a most important part in bringing about the present distribution of living things. From what has already been learned as to the past life of the various continents and their shifting connections with one another, it is now feasible to analyze the mammalian faunas of most of them and to separate the indigenous from the immigrant elements. Among the latter may be distinguished those forms which are the much modified descendants of ancient migrants from those which arrived at a much later date and have undergone but little change. To take a few examples from North America, it may be said that the Bears, Moose, Caribou and Bison are late migrants from the Old World; that the Virginia and Black-tailed Deer and the Prong-horned Antelope are of Old World origin, but their ancestors came in at a far earlier period and the modern species are greatly changed from the ancestral migrants. The Armadillo of Texas and the Canada Porcupine are almost the only survivors, north of Mexico, of the great migration of South American mammals which once invaded the northern continent. On the other hand, the raccoons and several families of rodents are instances of indigenous types which may be traced through a long American ancestry.