Fig. 122.—A gigantic Pampean †Ground-Sloth (†Megatherium americanum). Restored from a skeleton in the Museum of La Plata.
Opossums were extremely numerous, especially in the Brazilian caves, where in half a cubic foot of earth 400 jaws were collected.
The Pleistocene mammalian fauna of South America was a mixture of modern forms with ancient, vanished types similar to that which we found in North America. The †ground-sloths and †glyptodonts, the †litopterns, †toxodonts and †typotheres, the antelopes, horses and †mastodons have all disappeared from the continent, or vanished altogether from the face of the earth.
II. Tertiary Faunas
1. Pliocene
North America.—No part of the Cenozoic history of North America is so imperfectly recorded and so unsatisfactorily known as that of the Pliocene, and the later portion of that epoch is especially obscure. If the Peace Creek formation of Florida is properly referred to the upper Pliocene, it would show that the mammals of that time were substantially the same as those of the early Pleistocene.
The only fauna, as yet discovered, which can be referred to the middle Pliocene, is that of the Blanco beds of northwestern Texas, which have yielded but a scanty list of mostly ill-preserved fossils. Obviously, these give us a very incomplete picture of the life of that time. The great †ground-sloths had already reached North America, and the genus †Megalonyx, so common in the forested areas of Pleistocene North America, was perhaps already in existence. The †glyptodonts were likewise represented by one genus (†Glyptotherium) which was distinguished by the simple rings of the tail-sheath. No rodents have yet been found and only a few of the Carnivora, though a large cat, a musteline and a large “†bear-dog” are known. There were no true elephants, but several species of †mastodons, all of which were different from those of the Pleistocene; and in some, grinding teeth, though still low-crowned, had become much larger and more complex, marking a stage of advance toward the elephantine dentition. Horses of primitive type, the feet having three functional toes instead of one, were relatively abundant. Very large llama-like animals were present, but nothing has been ascertained with regard to the deer and antelopes of the time, and the only other representative of the Artiodactyla yet recovered is a peccary, interesting as being a species of the genus (†Platygonus) which became so abundant and widespread in the Pleistocene. Scanty and incomplete as this fauna is, it suffices to show that the middle Pliocene mammals were much more primitive than those of the Pleistocene.
Fig. 123.—†Horned Gopher (†Epigaulus hatcheri), lower Pliocene, Nebraska. Restored from a skeleton in the U.S. National Museum.