In the preceding chapter the origin of caves has been discussed, as well as their relation to the physical geography of the districts in which they are found. We must now pass on to the biological division of the subject, which relates to the animals that they contain and the inferences that may be drawn from their occurrence. The caves will be divided into historic, prehistoric, and pleistocene, according to the principles laid down in the [first] chapter.

It is extremely difficult, if not impossible, to define with precision the point where legend ends and history begins; but the line may be drawn with convenience at the first beginning of a connected and continuous narrative, rather than at the first isolated notice of a country. If we accept this definition, the historic period in Great Britain cannot be extended further back than the temporary invasion of Julius Cæsar, B.C. 55, even if so far, since of the interval that elapsed between that event and the subjugation under Claudius, in the year A.D. 43, we know scarcely anything. Of the events which happened in this country before Cæsar’s invasion there is no documentary evidence, although, by the modern method of scientific research, we are able to extend the narrative away from the borders of history far back into the archæological and geological past.

Wild Animals in Britain during the Historic Period.

During the historic period great changes have taken place in the animals inhabiting Great Britain. The wild animals have been diminished in number, and their area of occupation has been narrowed by the increase of population and the improvement in weapons of destruction. The brown bear, inhabiting Britain during the time of the Roman occupation, was extirpated probably before the tenth century. The current belief that it was destroyed in Scotland by the founder of the Gordon family in 1057 is unsupported by any documentary evidence which I have been able to discover; the crest of the Gordons, which is supposed to have been derived from the last of those animals slain in the island, consisting of three boars’, not bears’, heads. The last wolf is said to have been destroyed in Scotland in 1680, while in Ireland the animal lingered thirty years later to be a terror to the defenceless beggars. It was deemed worthy of a special decree for its destruction in the reign of Edward I. The wild boar was extinct before the reign of Charles I., while the beaver, which was hunted for its fur on the banks of the Teivi in Cardiganshire during the time of the first Crusade, became extinct shortly afterwards. The stag was so abundant in the south of England as recently as the reign of Queen Anne, that she saw a herd of no less than five hundred between London and Portsmouth. At present the animal lives only in a half-wild condition, in the forest of Exmoor and the Highlands of Scotland; while the roedeer is now only found wild in Scotland, although it formerly ranged throughout the length and breadth of the country.

The reindeer is proved to have been living in Caithness as late as the year 1159, by a passage in the Orkneyinga Saga.

The common rat, Mus decumanus, is the only wild or semi-wild animal that has migrated into this country during the historic period contrary to the will of man. In 1727 it (Pallas, Glires) had begun to invade Southern Russia from the regions of Persia and the Caspian Sea. Thence it swiftly spread over Asia Minor, and while it was advancing to the west overland, it was carried by ships to nearly all the ports in the world. It arrived in Britain certainly before the year 1730, and has since nearly exterminated the black indigenous species. It is the only wild animal which is known to have invaded Europe since the pleistocene age, with the exception, perhaps, of the true elk.

Animals living under the care of Man.

The fallow-deer, indigenous in the countries bordering on the Mediterranean, was probably introduced by the Romans, since its remains occur in refuse-heaps of Roman age, such as that of London Wall, and of Colchester, while it has not been met with in older deposits. To them, also, we probably owe the introduction of the pheasant, which was sufficiently abundant in the neighbourhood of London in the time of Harold to be mentioned as one of the articles of food eaten on feast-days by the households of the Canons at Waltham Abbey in 1059. The domestic fowl has left the first traces of its presence in this country in the Roman refuse-heaps, although it was known to the Belgæ, according to the testimony of Cæsar, before the first Roman invasion.

The earliest mention of the domestic cat in this country is to be found in the laws of Howel Dha,[40] that were probably codified at the end of the tenth or in the eleventh century, although many of the enactments may be of a much earlier date. The king’s cat is assessed at eightpence, or twice as much as that belonging to any subject. The ass[41] was certainly known in Britain in the days of Æthelred (A.D. 866–871), when, according to Professor Bell, its price was fixed at the large sum of twelve shillings. The larger breed of cattle represented by the Chillingham ox, and descended from the great Urus, first appears in this country about the time of the English invasion. It gradually spread over those districts conquered by the English, until the small aboriginal dark-coloured, short-horn Bos longifrons, which was the only domestic breed in the prehistoric and Roman times, is now only to be met with in the hill country of Wales and of Scotland, in which the Brit-Welsh or Romano-Celtic inhabitants still survive.

Classificatory value of Historic Animals.