Dinosaurs, with Special Reference to the American Museum Collections

by William Diller Matthew

Fig. 4.—Relative Length of Prehistoric and Historic Time.

There are indeed minor cycles of climate within this great cycle. The great Ice Age through which the earth has so recently passed was marked by alternations of severity and mildness of climate, of advance and recession of the glaciers, and within these smaller cycles are minor alternations whose effect upon the course of human history has been shown recently by Professor Huntington ("The Pulse of Asia"). But the great cycles of the geologic periods are of a scope far too vast for their changes to be perceptible to us except through their influence upon the course of evolution.

The Later Cycles of Geologic Time. The Reptilian Era opens with a period of extreme elevation, which rivalled that of the Glacial Epoch and was similarly accompanied by extensive glaciation of which some traces are preserved to our day in characteristic glacial boulders, ice scratches, and till, imbedded or inter-stratified in the strata of the Permian age. Between these two extremes of continental emergence, the Permian and the Pleistocene, we can trace six cycles of alternate submergence and elevation, as shown in the diagram (Fig. 5), representing the proportion of North America which is known to have been above water during the six geologic periods that intervene.

From this diagram it will appear that the six cycles or periods were by no means equal in the amount of overflow or complete recovery of the drowned lands. The Cretacic period was marked by a much more extensive and long continued flooding; the great plains west of the Mississippi were mostly under water from the Gulf of Mexico to the Arctic Ocean. The earlier overflows were neither so extensive nor so long continued. The great uplift of the close of the Cretacic regained permanently the great central region and united East and West, and the overflows of the Age of Mammals were mostly limited to the South Atlantic and Gulf coasts.

Sedimentary Formations. During the epochs of greatest overflow great marine formations were deposited over large areas of what is now dry land. These were followed as the land rose to sea level by extensive marsh and delta formations, and these in turn by scattered and fragmentary dry land deposits spread by rivers over their flood plains. In the marine formations are found the fossil remains of the sea-animals of the period; in the coast and delta formations are the remains of those which inhabited the marshes and forests of the coast regions; while the animals of the dryland, of plains and upland, left their remains in the river-plain formations.

Fig. 5.—Geologic Cycles and the Land Area of North America (after Schuchert).

These last, however, fragmentary and loose and overlying the rest, were the first to be swept away by erosion during the periods of elevation; and of such formations in the Age of Reptiles very little, if anything, seems to have been preserved to our day. Consequently we know very little about the upland animals of those times, if as seems very probable, they were more or less different from the animals of the coast-forests and swamps. The river-plain deposits of the Age of Mammals on the other hand, are still quite extensive, especially those of its later epochs, and afford a fairly complete record in some parts of the continent of the upland fauna of those regions.

Occurrence of Dinosaur Bones. Dinosaur bones are found mostly in the great delta formations, and since those were accumulated chiefly in the early stages of great continental elevations, it follows that our acquaintance with Dinosaurs is mostly limited to those living at certain epochs during the Age of Reptiles. In point of fact so far as explorations have yet gone in this country, the Dinosaur fauna of the close of the Jurassic and beginning of the Comanchic and that of the later Cretacic are the only ones we know much about. The immense interval of time that preceded, and the no less vast stretch of time that separated them, is represented in the record of Dinosaur history by a multitude of tracks and a few imperfect skeletons assigned to the close of the Triassic period, and by a few fragments from formations which may be intermediate in age between the Jurassic-Comanchic and the late Cretacic. Consequently we cannot expect to trace among the Dinosaurs, the gradual evolution of different races, as we can do among the quadrupeds of the Age of Mammals.