Fig. 2. Map showing locality records for Smilisca cyanosticta (triangles) and Smilisca phaeota (circles).
Hyla phaeota Cope, Proc. Acad. Nat. Sci. Philadelphia, 14 (9):358, 1862 [Holotype.—USNM 4347 from Turbo, Colombia; J. Cassin collector]. Boulenger, Catalogue Batrachia Salientia in British Museum, p. 402, Feb. 1, 1882. Werner, Sitzungsb. Akad. Wiss. München, 27:215, 1897. Günther, Biologia Centrali-Americana: Reptilia and Batrachia, p. 269, Sept. 1901. Nieden, Das Tierreich, Amphibia, Anura I, p. 261, June 1923. Dunn, Occas. Papers Boston Soc. Nat. Hist., 5:413, Oct. 10, 1931. Gaige, Hartweg, and Stuart, Occas. Papers Mus. Zool. Univ. Michigan, 357:4, Oct. 26, 1937. Cooper, Copeia, 2:122, June 30, 1944. Breder, Bull. Amer. Mus. Nat. Hist., 86(6):416, Aug. 26, 1946. Smith and Taylor, Bull. U.S. Natl. Mus., 194:88, June 17, 1948; Univ. Kansas Sci. Bull, 33:364, March 20, 1950. Taylor, Univ. Kansas Sci. Bull., 35(1):837, July 1, 1952. Brattstrom and Howell, Herpetologica, 10:117, [Pg 309] Aug. 1, 1954. Goin, Herpetologica, 14:120, July 23, 1958. Cochran, Bull. U.S. Natl. Mus., 220:57, 1961.
Hyla labialis Peters, Monats. Konigl. Akad. Wissen. Berlin, p. 463, 1863 [Holotype.—ZMB 4913 from "region of Bogotá," Colombia]; Monats. Konigl. Akad. Wissen. Berlin, p. 618, Oct. 16, 1873. Boulenger, Catalogue Batrachia and Salientia in British Museum, p. 397, Feb. 1, 1882.
Hyla baudini dolomedes Barbour, Occas. Papers Mus. Zool. Univ. Michigan, 129:11, Jan. 25, 1923 [Holotype.—MCZ 8539 from Río Esnápe, Sambú Valley, Darién, Panamá; Barbour and Brooks collectors]. Barbour and Loveridge, Bull. Mus. Comp. Zool. Harvard, 69:278, June, 1929.
Hyla phaeota phaeota, Smith, Herpetologica, 8:152, Jan. 30, 1953. Minton and Smith, Herpetologica, 16:103, June 17, 1960.
Smilisca phaeota, Starrett, Copeia, 4:303, Dec. 30, 1960.
Diagnosis.—Size large ([M] 65 mm., [F] 78 mm.); skull as long as wide, lacking frontoparietal fontanelle; large supraorbital flanges having straight edges and extending posterolaterally; large squamosal not in contact with maxillary; tarsal fold moderately wide, full length of tarsus; inner metatarsal tubercle moderately large, low, flat, elliptical; hind limbs relatively long, tibia averaging more than 53 per cent of snout-vent length; labial stripe silvery white; lips lacking vertical bars; loreal region pale green; dark brown or black tympanic mark dispersing into brown venated pattern on flanks; posterior surfaces of thighs pale brown, with or without darker flecks or small cream-colored flecks. (Foregoing combination of characters distinguishing S. phaeota from any other species in genus.)
| Table 2.—Geographic Variation in Size and Proportions in Males of Smilisca phaeota. (Means in Parentheses Below Observed Ranges; Data Based of Ten Specimens From Each Locality.) | |||
| Locality | Snout-vent length | Head width/ snout-vent length | Interorbital distance/ head width |
| Bonanza, Nicaragua | 40.8-47.7 | 34.1-38.0 | 31.0-36.1 |
| (43.7) | (36.3) | (35.4) | |
| Heredia Prov., Costa Rica | 46.3-59.0 | 32.5-36.0 | 30.5-39.6 |
| (51.7) | (35.0) | (34.7) | |
| Puntarenas Prov., Costa Rica | 53.6-64.9 | 32.6-36.1 | 31.0-38.0 |
| (61.4) | (34.5) | (34.4) | |
| Canal Zone, Panamá | 52.4-65.5 | 33.5-37.6 | 31.3-37.2 |
| (56.5) | (35.6) | (34.7) | |
| Río Quesada, Colombia | 52.6-61.0 | 33.1-37.1 | 30.1-33.9 |
| (56.0) | (35.0) | (32.1) | |