Ptychohyla euthysanota
Diagnosis.—Rostral keel absent; nuptial spines in males small; interorbital region much wider than eyelid.
Diagnosis.—Dorsum tan to reddish brown; venter white; rarely flecked with brown or black; a white stripe on upper lip, on flank, and usually above anus.
Description.—The following description is based on KU 58008 from Finca La Paz, Depto. San Marcos, Guatemala ([Pl. 12]). Adult male having a snout-vent length of 35.0 mm.; tibia length, 16.5 mm.; tibia length/snout-vent length, 47.1 per cent; foot length, 14.2 mm.; head length, 11.0 mm.; head length/snout-vent length, 31.4 per cent; head width, 10.7 mm.; head [width]/snout-vent length, 30.6 per cent; diameter of eye, 3.3 mm.; diameter of tympanum, 1.8 mm.; tympanum/eye, 54.5 per cent. Snout in lateral profile nearly square, slightly rounded above, and in dorsal profile bluntly rounded; canthus pronounced; loreal region moderately concave; lips thick, rounded, and slightly flaring; nostrils protuberant; internarial distance, 3.0 mm.; top of head flat; interorbital distance, 4.1 mm., and approximately a third broader than width of eyelid, 2.9 mm. Moderately heavy dermal fold from posterior corner of eye above tympanum to point above insertion of forelimb, covering upper edge of tympanum; tympanum round, its diameter slightly more than its distance from eye. Forearm moderately robust, having distinct dermal fold on wrist; dermal fold, but no row of tubercles along ventrolateral surface of forearm; pollex only slightly enlarged, bearing triangular shaped patch of small horn-covered spines (128 on right, 134 on left); second and fourth fingers equal in length; subarticular tubercles round, distal one on fourth finger bifid; discs moderate in size, that of third finger equal to diameter of tympanum; no web between first and second fingers; other fingers one-third webbed. Heels broadly overlap when hind limbs adpressed; tibiotarsal articulation reaches to middle of eye; low rounded tarsal fold; inner metatarsal tubercle large, elliptical, and flat; outer metatarsal tubercle small and round; low dermal fold from heel to disc of fifth toe; subarticular tubercles round; length of digits from shortest to longest 1-2-5-3-4; third and fifth toes webbed to base of disc; fourth toe webbed to proximal end of penultimate phalanx; thin dermal fold from inner metatarsal tubercle to disc of first toe; disc smaller than on fingers. Anal opening at the level of the upper edge of thighs; anal flap short; anal opening bordered above by thin transverse dermal fold and laterally by heavy dermal fold. Skin of dorsum and ventral surfaces of forelimbs and shanks smooth; that of throat, belly, and ventral surfaces of thighs granular. Ventrolateral glands moderately developed, not reaching axilla or groin and broadly separated midventrally. Tongue ovoid, emarginate, and only slightly free posteriorly; vomerine teeth 2-2, situated on small triangular elevations between ovoid inner nares; openings to vocal sac large, one situated along inner posterior edge of each mandibular ramus.
Dorsal ground-color of head, body, and limbs dull reddish brown with irregular dark brown reticulations on head and body and dark brown transverse bands on limbs; dorsal surfaces of first and second fingers and webbing on hand cream color; dorsal surfaces of third and fourth fingers dull brown; anterior surfaces of thighs dull creamy yellow; posterior surfaces of thighs dull brown; tarsi and toes tan with brown flecks; webbing of feet brown; faint creamy white stripe along lateral edges of tarsi and forearms; thin white line along edge of upper lip; distinct white stripe above and beside anal opening; axilla white; throat, chest, belly, and ventral surfaces of forelimbs creamy white; flanks white, separated from pale venter by a row of partly connected dark brown spots; ventral surfaces of thighs dull creamy yellow; feet grayish brown; ventrolateral glands pale grayish brown; small brown flecks on periphery of chin.
In life the dorsal ground-color was pale reddish brown (Orange-Cinnamon); dorsal reticulations dark brown (Chocolate); dorsal surfaces of first and second fingers and webbing on hands creamy tan (Light Pinkish Cinnamon); posterior surfaces of thighs reddish brown (Vinaceous-Tawny); webbing of feet gray (Deep Mouse Gray); throat and belly grayish white (Pale Gull Gray); ventral surfaces of hind limbs creamy white (Marguerite Yellow); spots on flanks dark brown (Warm Sepia); iris reddish bronze (Apricot Orange).
Variation.—No geographic variation in structural characters is discernible; variation in size and proportions is given in [Table 1]. Of 32 adults examined, seven have the tongue shallowly notched posteriorly; in the others the tongue is emarginate. Twenty specimens have a bifid subarticular tubercle beneath the fourth finger; in the others there are no bifid tubercles.