Life History.—This subspecies breeds in clear, swift mountain streams. Males call from stems and leaves of plants at the edge of, or overhanging, the streams. The breeding call consists of a soft "wraack," repeated at intervals of three to four seconds. Each note has a duration of 0.60 to 0.65 seconds and has 91 to 102 pulses per second; the dominant frequency falls between 3000 and 3200 cycles per second.

Tadpoles in various stages of development were found at Finca La Paz, Guatemala, in late July. This indicates that there is either extreme differential growth, or, more probably, an extended breeding season. A tadpole having a body length of 6.8 mm. and a total length of 19.1 mm. has a short median first upper tooth-row; lower tooth-rows 3-6 are only two-thirds as long as lower rows 1 and 2. Two recently metamorphosed young have snout-vent lengths of 14.2 and 14.8 mm.; they are colored like the adults.

Remarks.—The type specimen of Hyla euthysanota Kellogg (1928:123) is a female; therefore, when Taylor (1944) proposed the name Ptychohyla for hylids having ventrolateral glands in breeding males, he was unaware that Hyla euthysanota was a member of this group. In his description of Hyla rozellae, Taylor (1942) did not compare his specimens with Hyla euthysanota, but instead placed H. rozellae with H. loquax and H. rickardsi. The type series of H. rozellae consists of one large adult female and several metamorphosing young. Taylor (1949:16) based the description of Ptychohyla bogerti on two males and compared these specimens with P. adipoventris Taylor [= P. leonhardschultzei (Ahl)]. Thus, in a period of 22 years the females of this species were given two names and the male another. Stuart (1954:169) suggested that Hyla euthysanota and Hyla rozellae were Ptychohyla. Now that sufficient specimens are available from throughout the range it is possible to determine that the various named populations are conspecific.

Distribution.—This subspecies inhabits cloud forests at elevations of 660 to 2200 meters on the Pacific slopes of the Sierra Madre from extreme eastern Oaxaca and western Chiapas, México, through Guatemala to northern El Salvador; probably it occurs also in southwestern Honduras. Aside from the specimens listed below, three in the Frankfurt Museum from Depto. Santa Ana, El Salvador (44571, Hacienda San José; 43040, Hacienda Los Planes; 65119, Miramundo) are listed by Mertens (1952:29).

Specimens examined.—Mexico: Chiapas: Cascarada, 30 km. W of Cíltapec, UMMZ 87851-2; Cerro Ovando, UMMZ 87853-4; Chicomuselo, UMMZ 94439-40; Finca Juárez, 28 km. N of Escuintla, USNM 115052-5; Las Nubes, Cerro Ovando, USNM 115030-8; Salto de Agua, USNM 115039-51. Oaxaca: Cerro Pecho Blanco, UIMNH 40963; between La Gloria and Cerro Azul, UIMNH 40976-7; Río Grande, AMNH 51847-8; Santo Tomás Tecpan, UIMNH 41071.

Guatemala: San Marcos: Finca La Paz, 2 km. W of La Reforma, KU 58001-14, 59937 (skeleton), 60042-3 (tadpoles), 60044 (4 young), MCZ 34997, UMMZ 107739, 123151-7 (tadpoles); Finca Pirineos, Río Samalá, CNHM 35066. Santa Rosa: Finca La Gloria, UMMZ 123148 (tadpoles), 123150 (tadpoles). Sololá: Finca Santo Tomás, UMMZ 123149 (tadpoles); Olas de Mocá, near Mocá, CNHM 20208.

El Salvador: Chalatenango: Los Esemiles, USNM 73296. Santa Ana: Miramundo, CNHM 65120.

Plate 13

Click to View Larger. 		Ptychohyla euthysanota macrotympanum (Tanner) Hyla macrotympanum Tanner, Great Basin Nat., 17:52-53, July 31, 1957 [Holotype.—AMNH 62141 (formerly BYU 13752) from 10 miles east of Chiapa de Corzo, Chiapas, México; Robert Bohlman collector]. Ptychohyla macrotympanum, Duellman, Univ. Kansas Publ. Mus. Nat. Hist., 13:351, April 27, 1961.

Diagnosis.—Dorsum usually pale tan; venter white with scattered brown or black flecks; a thin white stripe on upper lip and another above anal opening; no distinct white stripe on flanks.