Fig. 312. Transverse Section of Segment of Tooth of Carboniferous Amphibian
On the other hand, the skull of the Carboniferous amphibians was made of close-set bony plates, like the skull of the reptile, rather than like that of the frog, with its open spaces (Figs. [313] and [314]). Unlike modern amphibians, with their slimy skin, the Carboniferous amphibians wore an armor of bony scales over the ventral surface and sometimes over the back as well.
It is interesting to notice from the footprints and skeletons of these earliest-known vertebrates of the land what was the primitive number of digits. The Carboniferous amphibians had five- toed feet, the primitive type of foot, from which their descendants of higher orders, with a smaller number of digits, have diverged.
The Carboniferous was the age of lycopods and amphibians, as the Devonian had been the age of rhizocarps and fishes.
| Fig. 313. Skull of a Permian Amphibian from Texas | Fig. 314. Skull of a Frog |
Life of the Permian. The close of the Paleozoic was, as we have seen, a time of marked physical changes. The upridging of the Appalachians had begun and a wide continental uplift—proved by the absence of Permian deposits over large areas where sedimentation had gone on before—opened new lands for settlement to hordes of air-breathing animals. Changes of climate compelled extensive migrations, and the fauna of different regions were thus brought into conflict. The Permian was a time of pronounced changes in plant and animal life, and a transitional period between two great eras. The somber forests of the earlier Carboniferous, with their gigantic club mosses, were now replaced by forests of cycads, tree ferns, and conifers. Even in the lower Permian the Lepidodendron and Sigillaria were very rare, and before the end of the epoch they and the Calamites also had become extinct. Gradually the antique types of the Paleozoic fauna died out, and in the Permian rocks are found the last survivors of the cystoid, the trilobite, and the eurypterid, and of many long-lived families of brachiopods, mollusks, and other invertebrates. The venerable Orthoceras and the Goniatite linger on through the epoch and into the first period of the succeeding era. Forerunners of the great ammonite family of cephalopod mollusks now appear. The antique forms of the earlier Carboniferous amphibians continue, but with many new genera and a marked increase in size.
A long forward step had now been taken in the evolution of the vertebrates. A new and higher type, the reptiles, had appeared, and in such numbers and variety are they found in the Permian strata that their advent may well have occurred in a still earlier epoch. It will be most convenient to describe the Permian reptiles along with their descendants of the Mesozoic.