Some peculiar branches of the camel stem appeared in North America. In the Pliocene arose a llama with the long neck and limbs of a giraffe, whose food was cropped from the leaves and branches of trees. Far more generalized in structure was the Oreodon, an animal related to the camels, but with distinct affinities also with other lines, such as those of the hog and deer. These curious creatures were much like the peccary in appearance, except for their long tails. In the middle Eocene they roamed in vast herds from Oregon to Kansas and Nebraska.
The ruminants. This division of the artiodactyls includes antelopes, deer, oxen, bison, sheep, and goats,—all of which belong to a common stock which took its rise in Europe in the upper Eocene from ancestral forms akin to those of the camels. In the Miocene the evolution of the two-toed artiodactyl foot was well-nigh completed. Bonelike growths appeared on the head, and the two groups of the ruminants became specialized,—the deer with bony antlers, shed and renewed each year, and the ruminants with hollow horns, whose two bony knobs upon the skull are covered with permanent, pointed, horny sheaths.
The ruminants evolved in the Old World, and it was not until the later Miocene that the ancestors of the antelope and of some deer found their way to North America. Mountain sheep and goats, the bison and most of the deer, did not arrive until after the close of the Tertiary, and sheep and oxen were introduced by man.
The hoofed mammals of the Tertiary included many offshoots from the main lines which we have traced. Among them were a number of genera of clumsy, ponderous brutes, some almost elephantine in their bulk.
The carnivores. The ancestral lines of the families of the flesh eaters—such as the cats (lions, tigers, etc.), the bears, the hyenas, and the dogs (including wolves and foxes)—converge in the creodonts of the early Eocene,—an order so generalized that it had affinities not only with the carnivores but also with the insect eaters, the marsupials, and the hoofed mammals as well. From these primitive flesh eaters, with small and simple brains, numerous small teeth, and plantigrade tread, the different families of the carnivores of the present have slowly evolved.
Dogs and bears. The dog family diverged from the creodonts late in the Eocene, and divided into two branches, one of which evolved the wolves and the other the foxes. An offshoot gave rise to the family of the bears, and so closely do these two families, now wide apart, approach as we trace them back in Tertiary times that the Amphicyon, a genus doglike in its teeth and bearlike in other structures, is referred by some to the dog and by others to the bear family. The well-known plantigrade tread of bears is a primitive characteristic which has survived from their creodont ancestry.
Cats. The family of the cats, the most highly specialized of all the carnivores, divided in the Tertiary into two main branches. One, the saber-tooth tigers ([Fig. 351]), which takes its name from their long, saberlike, sharp-edged upper canine teeth, evolved a succession of genera and species, among them some of the most destructive beasts of prey which ever scourged the earth. They were masters of the entire northern hemisphere during the middle Tertiary, but in Europe during the Pliocene they declined, from unknown causes, and gave place to the other branch of cats,—which includes the lions, tigers, and leopards. In the Americas the saber-tooth tigers long survived the epoch.
Fig. 351. Saber-Tooth Tiger