16. But one of the most obvious uses of our perceptions and our knowledge is to direct our Actions. It is suitable to the condition of our being that when we perceive a bunch of grapes, we should be able to pluck and eat the ripe ones; that when we perceive a stone rushing down the side of a hill, we should be able to move so as to avoid it. And this must be done by moving our limbs; in short, by the use of our muscles. And thus Sensation leads, not directly, but through the medium of Ideas, to muscular Contraction. I say that sensation and Muscular action are in such cases connected through the medium of Ideas. For when we proceed to pluck the grape which we see, the sensation does not determine the motion of the hand by any necessary geometrical or mechanical conditions, as an impression made upon a machine determines its motions; but the perception leads us to stretch forth the hand to that part of space, wherever it is, where we know that the grape is; and this, not in any determinate path, but, it may be, avoiding or removing intervening obstacles, which we also perceive. There is in every such case a connexion between the sensation and the resulting action, not of a material but of a mental kind. The cause and the effect are bound together, not by physical but by intellectual ties.
17. And thus in such cases, between the two vital operations, Sensation and Muscular Action, there intervenes, as an intermediate step, Perception or Knowledge, which is not merely vital but ideal. But this is not all; there is still another mental part of the process which may be readily distinguished from that which we have described. An act of the Will, a Volition, is that, in the Mind, which immediately determines the action of the Muscles of the Body. And thus Will intervenes between Knowledge and Action; and the cycle of operations which take place when animals act with reference to external objects is [234] this:—Sensation, Perception, Volition, Muscular Contraction.
18. To attempt further to analyse the mental part of this cycle does not belong to the present part of our work. But we may remark here, as we have already remarked in the History[97], how irresistibly we are led by physiological researches into the domain of thought and mind. We pass from the body to the soul, from physics to metaphysics; from biology to psychology; from things to persons; from nouns to pronouns. I have there noticed the manner in which Cuvier expresses this transition by the introduction of the pronoun: ‘The impression of external objects upon the me, the production of a sensation, of an image, is a mystery impenetrable to our thoughts.’
[97] Hist. Ind. Sc. b. xvii. c. v. s. 2.
19. But to return to the merely biological part of our speculations. We have arrived, it will be perceived, at this result: that in animal actions there intervenes between the two terms of Sensation and Muscular Contraction, an intermediate process; which may be described as a communication to and from a Center. The Center is the seat of the sentient and volent faculties, and is of a hyperphysical nature. But the existence of such a Center as a necessary element in the functions of the animal life is a truth which is important in biology. This indeed may be taken as the peculiar character of animal, as distinguished from merely organic powers. Accordingly, it is so stated by Bichat. For although he superfluously, as I have tried to show, introduces into his list of vital powers an organic sensibility, he still draws the distinction of which I have spoken; ‘in the animal life, Sensibility is the faculty of receiving an Impression plus that of referring it to a common Center[98].’
[98] Life and Death, p. 84.
20. But since Sensibility and Contractility are thus connected by reference to a common Center, we may ask, before quitting the subject, what are the different forms which this reference assumes? Is the connexion [235] always attended by the distinct steps of Knowledge and Will,—by a clear act of consciousness, as in the case which we have taken, of plucking a grape; or may these steps become obscure, or vanish altogether?
We need not further illustrate the conscious connexion. Such actions as we have described are called voluntary actions. In extreme cases, the mental part of the process is obvious enough. But we may gradually pass from these to cases in which the mental operation is more and more obscure.
In walking, in speaking, in eating, in breathing, our muscular exertions are directed by our sensations and perceptions: yet in such processes, how dimly are we conscious of perceptive and directive power! How the mind should be able to exercise such a power, and yet should be scarcely or not at all conscious of its exercise, is a very curious problem. But in all or in most of the instances just mentioned, the solution of this problem appears to depend upon psychological rather than biological principles, and therefore does not belong to this place.
21. But in cases at the other extreme (unconscious actions) the mental part of the operation vanishes altogether. In many animals, even after decapitation, the limbs shrink when irritated. The motions of the iris are determined by the influence of light on our eyes, without our being aware of the motions. Here Sensations produce Motions, but with no trace of intervening Perception or Will. The Sensation appears to be reflected back from the central element of animal life, in the form of a Muscular Contraction; but in this case the Sensation is not modified or regulated by any Idea. These reflected motions have no reference to relations of space or force among surrounding objects. They are blind and involuntary, like the movements of convulsion, depending for direction and amount only on the position and circumstances of the limb itself with its muscles. Here the Centre from which the reflection takes place is merely animal, not intellectual.