[106] Anatomie Générale, i. liii.

12. This Idea of a Final Cause is applicable as a fundamental and regulative idea to our speculations concerning organized creatures only. That there is a purpose in many other parts of the creation, we find abundant reason to believe, from the arrangements and laws which prevail around us. But this persuasion is not to be allowed to regulate and direct our reasonings with regard to inorganic matter, of which conception the relation of means and end forms no essential part. In mere Physics, Final Causes, as Bacon has observed, are not to be admitted as a principle of reasoning. But in the organical sciences, the assumption of design and purpose in every part of every whole, that is, the pervading idea of Final Cause, is the basis of sound reasoning and the source of true doctrine.

13. The Idea of Final Cause, of end, purpose, design, intention, is altogether different from the Idea of Cause, as Efficient Cause, which we formerly had to consider; and on this account the use of the word Cause in this phrase has been objected to. If the idea be clearly entertained and steadily applied, the word is a question of subordinate importance. The term Final Cause has been long familiarly used, and appears not likely to lead to confusion. [248]

14. The consideration of Final Causes, both in physiology and in other subjects, has at all times attracted much attention, in consequence of its bearing upon the belief of an Intelligent Author of the Universe. I do not intend, in this place, to pursue the subject far in this view: but there is one antithesis of opinion, already noticed in the History of Physiology, on which I will again make a few remarks[107].

[107] Hist. Ind. Sc. b. xvii. c. viii. On the Doctrine of Final Causes in Physiology.

It has appeared to some persons that the mere aspect of order and symmetry in the works of nature—the contemplation of comprehensive and consistent law—is sufficient to lead us to the conception of a design and intelligence producing the order and carrying into effect the law. Without here attempting to decide whether this is true, we may discern, after what has been said, that the conception of Design, arrived at in this manner, is altogether different from that Idea of Design which is suggested to us by organized bodies, and which we describe as the doctrine of Final Causes. The regular form of a crystal, whatever beautiful symmetry it may exhibit, whatever general laws it may exemplify, does not prove design in the same manner in which design is proved by the provisions for the preservation and growth of the seeds of plants, and of the young of animals. The law of universal gravitation, however wide and simple, does not impress us with the belief of a purpose, as does that propensity by which the two sexes of each animal are brought together. If it could be shown that the symmetrical structure of a flower results from laws of the same kind as those which determine the regular forms of crystals, or the motions of the planets, the discovery might be very striking and important, but it would not at all come under our idea of Final Cause.

15. Accordingly, there have been, in modern times, two different schools of physiologists, the one proceeding upon the idea of Final Causes, the other school [249] seeking in the realm of organized bodies wide laws and analogies from which that idea is excluded. All the great biologists of preceding times, and some of the greatest of modern times, have belonged to the former school; and especially Cuvier, who may be considered as the head of it. It was solely by the assiduous application of this principle of Final Cause, as he himself constantly declared, that he was enabled to make the discoveries which have rendered his name so illustrious, and which contain a far larger portion of important anatomical and biological truth than it ever before fell to the lot of one man to contribute to the science.

The opinions which have been put in opposition to the principle of Final Causes have, for the most part, been stated vaguely and ambiguously. Among the most definite of such principles, is that which, in the History of the subject, I have termed the Principle of Metamorphosed and Developed Symmetry, upon which has been founded the science of Morphology.

The reality and importance of this principle are not to be denied by us: we have shown how they are proved by its application in various sciences, and especially in botany. But those advocates of this principle who have placed it in antithesis to the doctrine of Final Causes, have, by this means, done far more injustice to their own favourite doctrine than damage to the one which they opposed. The adaptation of the bones of the skeleton to the muscles, the provision of fulcrums, projecting processes, channels, so that the motions and forces shall be such as the needs of life require, cannot possibly become less striking and convincing, from any discovery of general analogies of one animal frame with another, or of laws connecting the development of different parts. Whenever such laws are discovered, we can only consider them as the means of producing that adaptation which we so much admire. Our conviction that the Artist works intelligently, is not destroyed, though it may be modified and transferred, when we obtain a sight of his tools. Our discovery of laws cannot contradict our persuasion of ends; our Morphology cannot prejudice our Teleology. [250]

16. The irresistible and constant apprehension of a purpose in the forms and functions of animals has introduced into the writings of speculators on these subjects various forms of expression, more or less precise, more or less figurative; as, that ‘animals are framed with a view to the part which they have to play;’—that ‘nature does nothing in vain;’ that ‘she employs the best means for her ends;’ and the like. However metaphorical or inexact any of these phrases may be in particular, yet taken altogether, they convey, clearly and definitely enough to preclude any serious errour, a principle of the most profound reality and of the highest importance in the organical sciences. But some adherents of the morphological school of which 1 have spoken reject, and even ridicule, all such modes of expression. ‘I know nothing,’ says M. Geoffroy Saint Hilaire, ‘of animals which have to play a part in nature. I cannot make of nature an intelligent being who does nothing in vain; who acts by the shortest mode; who does all for the best.’ The philosophers of this school, therefore, do not, it would seem, feel any of the admiration which is irresistibly excited in all the rest of mankind at the contemplation of the various and wonderful adaptations for the preservation, the enjoyment, the continuation of the creatures which people the globe;—at the survey of the mechanical contrivances, the chemical agencies, the prospective arrangements, the compensations, the minute adaptations, the comprehensive interdependencies, which zoology and physiology have brought into view, more and more, the further their researches have been carried. Yet the clear and deep-seated conviction of the reality of these provisions, which the study of anatomy produces in its most profound and accurate cultivators, cannot be shaken by any objections to the metaphors or terms in which this conviction is clothed. In regard to the Idea of a Purpose in organization, as in regard to any other idea, we cannot fully express our meaning by phrases borrowed from any extraneous source; but that impossibility arises precisely from the circumstance of its being a Fundamental Idea which is inevitably assumed in our [251] representation of each special fact. The same objection has been made to the idea of mechanical force, on account of its being often expressed in metaphorical language; for writers have spoken of an energy, effort, or solicitation to motion; and bodies have been said to be animated by a force. Such language, it has been urged, implies volition, and the act of animated beings. But the idea of Force as distinct from mere motion,—as the Cause of motion, or of tendency to motion,—is not on that account less real. We endeavour in vain to conduct our mechanical reasonings without the aid of this idea, and must express it as we can. Just as little can we reason concerning organized beings without assuming that each part has its function, each function its purpose; and so far as our phrases imply this, they will not mislead us, however inexact, or however figurative they be.