c. The Prostate Gland.—That the prostate gland is intimately associated with reproduction is evident from the fact that in those male animals that have suffered castration before puberty, the prostate gland withers and practically disappears. What then is the role that this gland plays? Like Cowper's glands, it secretes only during sexual excitement. Under such excitement its ducts become gorged with a secretion peculiar to it and at the moment of the emission or the ejaculation of the semen the numerous ducts empty their contents into the urethra to be mingled with and made a part of the semen.

The secretion of the prostate is composed of a watery solution of protein and of alkaline salts and so closely similar to the secretion of the seminal vesicles that we will consider its action along with that of the secretion from the vesicles.

d. The Seminal Vesicles.The seminal vesicles secrete continuously. The secretion is composed of an aqueous solution of albumin and of alkaline salts. This secretion together with the secretion from the prostate gland is poured into the urethra at the moment of sexual orgasm; they become mixed in their transit through the urethra with the secretion from the testes. This mixture is known as semen. [See pg. 42.]

It used to be supposed that the semen was secreted wholly by the testes; that the testes were secreting continuously and that the seminal vesicles were receptacles for the gradually accumulating semen from the testes. The researches of Steinach and others have made the old theory untenable and demonstrate that the semen is a mixture from three distinct sources; that the testicles secrete their contribution to the semen only during sexual stimulation; while the seminal vesicles secreting their products continuously become periodically filled and distended.

Let us inquire regarding the function of this alkaline albuminous secretion from the vesicles and prostate. For what purpose does Nature prepare such a secretion? The spermatozoa frequently remain several days in the organs of the female before the ovum is found and fertilized. During these several days the spermatozoa are exerting no small amount of energy in their vigorous flagellate movement. For such an expenditure of energy they must receive nourishment and stimulation. The nourishment is supplied by the albumin and proteid of the vesicular and prostatic secretions. The stimulation is supplied by the salts also secreted by these glands. The recent researches of Loeb and others have demonstrated the importance of mineral salts in stimulating the activity of living cells. One can cite no better example of this stimulant action than the influence of these vesicular and prostatic salts upon the activity of the spermatozoa.

The vesicles and prostate may be looked upon as the commissariat of the army of spermatozoa; the vesicles accumulating a stock of supplies to be drawn upon at short notice; the prostate representing a factory where a considerable quantity of supplies can be prepared at short notice.

This periodic distention of the seminal vesicles is a matter of very considerable hygienic importance and must be thoroughly understood by every young man who would lead a normal sexual life.

These organs in common with all other organs of the body are supplied with two sets of nerves, one set passing away to the spinal cord and carrying messages which indicate the condition of the organ or the presence and character of any local stimulus; the other passing away from the spinal cord to the organ and carrying secretory and motor impulses. The secretory impulses are more or less continuous and as a result, these glands secret continuously and become periodically distended as described above. The motor impulses pass to the muscles within the walls of the vesicles, causing a strong spasmodic contraction of these muscles at the moment of emission of semen, thus throwing the contents of the vesicles into the urethra at the same moment when the epididymis the vas deferens and the ducts of the prostate are emptying their secreted contents into the urethra.

Now the sensory nerves passing from the seminal vesicles up to the erection and emission centers are stimulated by any unusual pressure within the vesicles. Unusual pressure may be caused either by distention due to accumulated secretion or by pressure upon the vesicles from over-distended rectum or bladder. It sometimes happens that two or more of these influences are acting at the same time. These impulses are most likely to be effective when the subject is asleep, and particularly if he is lying upon his back. The result of the stimulus is to cause an erection, accompanied usually by an erotic dream, the whole phenomenon culminating in an emission of the contents of the seminal vesicles and followed, of course, by a relief of the pressure which was the cause of the condition. This phenomenon has been variously called nocturnal emission, "pollution" and "dreaming-off."

Vecki, a specialist in physiology, hygiene and pathology of the sexual apparatus, says that the nocturnal emission is a normal physiological phenomenon, the object of which is to relieve pressure in the seminal vesicles, and that in normal cases it occurs in fairly regular periods, these periods varying in length with different individuals, according to their physical condition and habits, the period being two to four weeks, usually; though a considerably longer or shorter period would not be looked upon as pathological. Vecki describes the normal nocturnal emission as being accompanied by an erection, erotic dreams, and an orgasm, the subject being wholly unconscious of the condition until he is awakened at the moment of orgasm. Normally, the subject experiences on the following day a feeling of relief and well-being and should, normally, be wholly free from headache, depression or languor.