Of the Venous Pulse.
The pulsation that the veins have under certain circumstances, must not be taken for an effect of the venous irritability. It is an effect of the reflux of the blood, which not being able to go through the lungs, stagnates in the pulmonary arteries and in the right side of the heart; so that when this contracts, as the blood finds an obstacle in the ordinary course, it flows back whence it came, as when the aliments are unable to pass down, they take the other direction. This reflux takes place, to a certain distance, notwithstanding the valves; it is often very evident in the jugular vein, when animals, submitted to experiments, breathe laboriously; then it is discontinued; it takes place three or four times, then ceases, and returns irregularly; it is observed also in the last moments of life, when the lungs are embarrassed.
The vein is then sensibly dilated; then it contracts. But if you apply the finger above, you do not experience a sensation analogous to that of the pulse; you will perceive only a wave of blood which flows back. The reason of it is plain; 1st, there is no locomotion; 2d, as the venous parietes are loose, they could not strike the finger sufficiently strong, if there was a similar change of place. Observe, that it is less the blood than the artery itself which by its firm texture gives the sensation of the pulse; if it could straighten itself when empty, as it does when it is full, it would produce nearly the same sensation; this is a remark that should be added to what I have said upon the pulse in the preceding system.
The contraction of the veins in the motion of the reflux, of which we are treating, is only the contractility of texture. When the heart ceases to propel the blood in its cavity, it contracts, after having been dilated; it is nearly the same in the dead body, in which we fix a syringe in the veins; when they are very full of water, if we draw back the piston a little, immediately the fluid returning, the vein contracts; it is as when it contracts from a puncture that evacuates the blood; this does not imply any irritability.
I believe that sometimes this reflux may depend upon an irregular motion of the heart, which contracts in an opposite direction to the ordinary one, though there is no obstacle in the lungs. What induces me to think so, is, that frequently in experiments, at the moment the animal begins to suffer much, the reflux takes place before the lungs have had time to be disturbed. A very remarkable thing in experiments is the quickness with which pain disturbs the motion of the heart, accelerates it, renders it irregular, &c. We can always at will hasten respiration, by making the animal suffer; now the acceleration of the pulse is always prior to that of respiration, which appears to be determined by it. I am persuaded that if the diseases of the heart were as frequent on the right side as the left, they would often produce this reflux and this pulsation of the veins.
The limits of the reflux of the venous blood vary. Haller has observed it as far as the iliacs. In general, it rarely goes beyond the great trunks, on account of the valves. I have demonstrated in my Researches upon Death, that the colour of those who die of asphyxia, of those who are drowned, &c. does not depend on this, because it cannot evidently extend to the capillary system, which receives the black blood that colours it, from the arteries that then circulate that kind of blood.
The reflux of the black blood in the veins, produced in the preceding cases, either by an obstruction in the lungs, or by a sudden derangement in the action of the heart, takes place in a natural state, though in an infinitely less degree. In fact, when the right auricle contracts, all the blood does not pass into the corresponding ventricle; the veins being open, a portion flows back into them. It is difficult to determine the extent of this natural reflux, of which all authors have spoken. When the thorax is opened, we observe it distinctly; we might then ascertain its extent; but in this case, respiration not being performed as usual, it is evident that we cannot judge by it of what ordinarily takes place.
Insensible Contractility.
This property, which, like the preceding, is inseparable from the organic sensibility, exists in the veins as in the other parts; it presides only over nutrition; it appears more evident than in the arteries; at least the diseases which particularly increase it are more frequent in the veins. The texture of these vessels is often inflamed. 1st. Bell relates cases of it, the effect of external violence. 2d. Every one is acquainted with the inflammation of the hemorrhoids. 3d. The cicatrization of venous wounds after bleeding is a product of inflammation. Without doubt this cicatrization is promoted by the want of impulse, to which the arteries are subjected; but certainly these last would not in like circumstances heal so fast, if they did at all. When an artery has been tied, it is necessary that its parietes, inflamed by the action of the thread, most often cut by it, and brought into contact, should form adhesions, that the cure may be complete, and the ligature come off without danger. Now, nothing is more difficult and slower than their adhesion, from the difficulty with which the arterial texture inflames. Hence the frequency of hemorrhage after the operation for aneurism and other great operations. The blood often bursts out at the end of twenty, thirty, or forty days; the surgeon should always be upon his guard when he has tied these great trunks, from the want of disposition in the arterial texture to inflame. Frequently when the artery is obliterated, it is not by inflammation. Whilst the ligature stops the blood, the portion of artery comprised between it and the first collateral branch, closes gradually by the contractility of texture, and forms a kind of ligament, which arrests the blood after the thread has fallen off. I do not know but that these cases are more numerous than those of inflammation. Now the veins always adhere soon when they are tied; their wounds cicatrize immediately. In great wounds it is almost always useless to tie them at the first moment, on account of the valves, as I have said above, and afterwards, because the cut ends contract, and soon inflame and adhere. If there are venous hemorrhages, it is at the time of the injury, and not as long after as in the arteries.
Every thing proves, then, that the vital activity is much greater in the venous than in the arterial system, in respect to the tonic powers. The absence of the cellular texture in the second and its presence in the first, may have an influence upon this phenomenon.