2d. The internal absorptions, on the contrary, every where correspond with the analogous exhalations. Thus the absorbents take up, upon the serous system, the serum, upon the cellular system the serum and fat, upon the medullary system the marrow, upon the synovial system, the synovia; fluids, all of which as we have seen, had been thrown out by exhalation upon their respective surfaces, and had remained for an instant upon them. These absorptions are made in a constant and regular manner; it is in this that they differ from the preceding. The internal absorbents, incessantly in action, take up in a given time the same quantity of fluids; their action corresponds precisely with that of the exhalants. Observe that there are two essential differences between the external and internal absorptions; viz. that the one are exerted on the one hand on fluids different from those exhaled upon their surfaces, and that on the other they are subject to continual variations and irregularities; whilst the others always take up the fluids exhaled upon their surfaces, and are constant and regular, at least in a state of health. I shall point out in the mucous and cutaneous, systems, the cause of this important difference.
3d. As to the nutritive absorptions, we know much less of them than the preceding; but nutrition evidently supposes them. There is in fact in this function a double motion, one of composition, the other of decomposition. No organ, no part of an organ is formed at one period of the same elements that composed it at a former one. The ancients thought, without positive proof, that the body was renewed every seven years. Whatever may be the period of its renewal, we cannot deny that it is continually composed and decomposed; now the exhalants perform the first nutritive motion, the absorbents the second. Observe in fact, that the internal substances never re-enter the circulation to be afterwards thrown out, except by the way of the absorbents.
The nutritive absorptions differ then from the preceding, in this, that the substance deposited by exhalation and taken up by them, remains in the organs, makes a part of them, and contributes to their composition; whilst the fluids with which the internal exhalations and absorptions are concerned, after having been furnished by the one, and before being taken up by the other, remain out of the organs, upon their surface or in their cells, but do not make a part of their structure.
It is difficult perhaps to conceive how solid nutritive substances can be absorbed by vessels so delicate. Hunter, to whom anatomy owes much both as it respects the absorbents and their uses, has already removed this objection. It may be added to what he has said, that the distinction between solids and fluids is not substantial except when they are in mass; but when they are considered as separate particles, they do not differ; this is so true, that the same particle makes alternately part of a solid and a fluid, as in water that is not frozen and that which is, as in solid or melted lead, &c. Now the nutritive substances are absorbed particle by particle; then the distinction of fluid and solid is of no consequence in the function of absorption.
Since the origin of the absorbents is beyond the reach of our senses, it is difficult, it is impossible even to determine the manner in which they arise, the peculiar structure which distinguishes them at their origin, their communications, &c. They undoubtedly differ essentially according to the mucous, cutaneous, serous, synovial, cellular, medullary surfaces to which they belong; undoubtedly the nutritive absorbents differ remarkably from the others; but nothing can be proved by inspection. What has not been said upon the villous coat of the intestines considered as the origin of the lacteals, upon their small bladders, upon the form of the pores of the peritoneum, the pleura, &c. upon the cellular sponge? I shall not notice here these anatomical hypotheses, which have been made by an abuse of the microscope, and which besides, if they had any real foundation, would not lead to any inference useful to science.
Do the absorbents arise from the capillary system? Judging by injections, it would seem that they did, for many distinguished anatomists, by forcing a fine injection through the arteries, have filled the absorbents in the neighbourhood. I never saw any thing similar to it myself, yet I am far from denying a fact attested by Meckel. If many other experiments should confirm it, it would be established incontestably, it is evident, that the origin of the absorbents is in the capillary system, as the origin of the excretories and exhalants are proved to be in the same system. Besides, the phenomena of absorptions cannot give us any light upon the mode of the origin of the absorbents.
Where they go off from the surfaces or the organs from which they arise, the absorbents are extremely delicate; they elude all kinds of injection. They appear to anastomose with each other, interlace, form a complicated net-work, which contributes much to the structure of some parts, especially of the serous membranes. We know however but little of this interlacing. It is not until they have run a certain course, that these vessels are cognizable by our senses, and that we can consequently study them in a general manner.
II. Course of the Absorbents.
The absorbents, arising from the different parts that we have pointed out, go in different ways.
1st. In the extremities, they are divided immediately into two very distinct courses, the one superficial, the other deep-seated. The former accompanies at first the sub-cutaneous veins, then runs along in their interstices; so that when injections have succeeded well, the whole exterior of the limbs appears to be covered with a kind of lymphatic net-work. The second goes along the muscular interstices, principally in the course of the arteries and the veins. Both tend towards the superior parts of the limbs. When the vessels arrive there, they approximate each other, and are collected into a bundle, in which they are fewer but larger than below, and which passes through certain openings that lead them into the trunk; for example, those of the superior extremities almost all terminate in the axilla, those of the inferior in the groin and some in the ischiatic notch. Now as it is a general rule, that every absorbent should pass through one or more glands, nature has placed at these openings of communication of the extremities with the trunk, a certain number of these glands. Yet before arriving at them, some have already passed through similar glands placed, in a less number, it is true, in the ham and the bend of the arm. It is in the extremities that the absorbents run the longest course without passing through glands.