1. SAMBUCUS L. Elder.
Trees or shrubs, with stout branches containing thick white or brown pith, and buds with several scales. Leaves petiolate, unequally pinnate, deciduous, with serrate or laciniate leaflets, the base of the petiole naked, glandular or furnished with a stipule-like leaflet; stipels small, leaf-like, usually setaceous, often 0; stipules small, rudimentary, usually 0 except on vigorous shoots. Flowers small, in broad terminal corymbose cymes, their bracts and bractlets lanceolate, acute, scarious, caducous, sometimes ebracteolate; calyx-tube ovoid, the limb 3—5-lobed or toothed; corolla rotate or slightly campanulate, equally 3—5-parted; filaments filiform or subulate; ovary inferior or partly superior, 3—5-celled; style abbreviated, thick and conic, 3—5-lobed, stigmatic at apex. Fruit subglobose, with juicy flesh, and 3—5 oblong cartilaginous punctate-rugulose or smooth 1-seeded nutlets full and rounded on the back and rounded at the ends. Seeds filling the cavity of the nutlets, pale brown; cotyledons ovoid.
Sambucus with about twenty species is widely and generally distributed through the temperate parts of North America, Europe, and Asia, and inhabits high mountain ranges within the tropics, and in Australia, Tasmania, and New Zealand. Of the nine or ten North American species three are arborescent. Sambucus possesses cathartic and emetic properties in the bark; the flowers are excitant and sudorific, and the juice of the fruit is alterative and laxative. The dried flowers of the European Sambucus nigra L., are used in the preparation of an aromatic distilled water and in flavoring lard, and the hard and compact wood is made into combs and mathematical instruments. The large pithy shoots of Sambucus furnish children with pop-guns, pipes, and whistles; and the fruit of some of the species is cooked and eaten.
Sambucus, the name of the Elder-tree, is believed to have been derived from σαμβύκη, a musical instrument, probably in allusion to the use of the pithy stems.
CONSPECTUS OF THE ARBORESCENT SPECIES OF THE UNITED STATES.
Cymes flat-topped; pith usually white; fruit black; nutlets rugose. Fruit lustrous.1. [S. Simpsonii] (C). Fruit appearing blue from a thick covering of bloom.2. [S. coerulea] (B, F, G, H). Cymes ovoid; pith pale brown; fruit red; nutlets smooth.3. [S. callicarpa] (B, G).
1. [Sambucus Simpsonii] Rehd.
Leaves 4′—7′ long, 3—7, usually 5-foliolulate, with a glabrous petiole and usually 5 dark yellow-green leaflets, lustrous and glabrous on the upper surface with the exception of a few scattered hairs on the midrib, and paler and glabrous on the lower surface, the terminal leaflet obovate or oblong-obovate, short-acuminate at apex, and gradually narrowed at base into a slender petiolule ⅓′—½′ in length, the lateral leaflets broad-elliptic to oblong-elliptic, short-acuminate, broad-cuneate at base, those of the upper pair usually sessile, those of the lower pair on short stalks rarely more than 1/12′ long, serrate except at the base with small slightly spreading teeth, 1½′—3′ long and 1½′—2½′ wide. Flowers slightly fragrant, on slender pedicels in convex or sometimes flat cymes 3′—8′ in diameter, with 4 or 5 rays, the terminal ray as long or longer than the lateral rays, rarely shorter; calyx-tube ovoid, the lobes oblong-ovate, acute, about as long as the tube and slightly exceeding the thick conic style; stamens about as long as the white corolla-lobes; ovary usually 5, rarely 4-celled. Fruit subglobose, dark purplish black, about ¼′ in diameter; nutlets rugose.
A tree, sometimes 15°—18° high, with a trunk often 8′ in diameter, and slightly angled branchlets greenish when they first appear, becoming light yellow-gray and sometimes covered during their second and third years with thick corky excrescences; pith white, on 2 or 3-year-old branches comparatively narrow, occupying only about one-third of the diameter of the stem.
Distribution. Florida, neighborhood of Jacksonville, Duval County, to Eustis, Lake County, Bradentown, Manatee County, and Sanibel Island, Lee County; Mississippi, Ocean Springs, Jackson County; Louisiana, Cameron, Cameron Parish.
2. [Sambucus coerulea] Raf.
Sambucus glauca Nutt.
Sambucus neomexicana Woot.
Leaves 5′—7′ long, with a stout grooved petiole much enlarged and naked or sometimes furnished at the base with leaf-like appendages, and 5—9 ovate or narrow-oblong leaflets contracted at apex into a long point, unequally cuneate or rounded at base, and coarsely serrate with spreading or slightly incurved callous-tipped teeth, the lower leaflets often 3-parted or pinnate, the terminal one sometimes furnished with 1 or 2 lateral stalked leaflets, yellow-green on the upper surface, pale on the lower surface, covered with scattered pale hairs when they unfold, and at maturity glabrous or soft pubescent beneath (var. velutina Rehd.), thin, rather firm in texture, bright green above and pale below, 1′—6′ long and ⅓′—1½′ wide, with a narrow pale midrib and inconspicuous veins; petiolules slender, those of the lateral leaflets ¼′—½′ and of the terminal leaflet up to 2′ in length; stipels linear, oblong-lanceolate to ovate, rounded or acute at apex, entire or sharply serrate and leaf-like, 1/16′—½′ long, caducous, often 0. Flowers ⅛′ in diameter, appearing from April in southern California to July in British Columbia, in flat long-branched glabrous or pubescent cymes 4′—10′ wide, with linear acute green caducous bracts and bractlets, the lower branches often from the axils of upper leaves; flower-buds globose, covered with a glaucous bloom, sometimes turning red before opening; calyx ovoid, red-brown, with acute scarious lobes; corolla yellowish white, with oblong divisions rounded at apex, as long as the stamens. Fruit subglobose, ⅓′ in diameter, black, appearing blue by its thick covering of mealy bloom; flesh rather sweet and juicy.
A tree, 30°—50° high, with a tall straight trunk sometimes enlarged at base and 12′—18′ in diameter, stout spreading branches forming a compact round-topped head, and branchlets usually without a terminal bud, green tinged with red or brown when they first appear, and covered with short white caducous hairs, or densely soft pubescent during their first season (var. velutina Rehd.), stout, slightly angled, covered with lustrous red-brown bark in their first winter and nearly encircled by the large triangular leaf-scars marked by conspicuous fibro-vascular bundle-scars; pith white or rarely brownish; often a broad shrub, with numerous spreading stems. Winter-buds axillary generally in pairs, superposed or in clusters of 4 or 5, only the upper bud or sometimes the lower usually developing, covered with 2 or 3 pairs of opposite broad-ovate chestnut-brown scales, those of the inner rank accrescent, and at maturity acute, entire, green, 1′ long, and sometimes developing into pinnate leaves 2′—3′ in length. Bark of the trunk deeply and irregularly fissured, the dark brown surface slightly tinged with red and broken into small square appressed scales. Wood light, soft, weak, coarse-grained, yellow tinged with brown, with thin lighter colored sapwood.
Distribution. Gravelly rather dry soil of valleys and river-bottoms; western Montana (neighborhood of Flathead Lake and Missoula, Missoula County), through Idaho to the coast of British Columbia (Vancouver Island), and southward to the San Bernardino Mountains and Santa Catalina Island, California, ascending on the Cascade and Sierra Nevada Mountains to altitudes of 6000°—8000°; Nevada, King’s Cañon, Ormsby County; Utah, Juab, Juab County, and the neighborhood of Salt Lake City, Salt Lake County; Colorado, near Trinidad, Las Animas County; New Mexico, Sacramento Mountains, Otero County; very abundant in the coast region; comparatively rare in the interior; of its largest size in the valleys of western Oregon; northward, and east of the Cascade and Sierra Nevada Mountains rarely arborescent; in southern California often with smaller leaves and flower-clusters than northward; the var. velutina rare and local, California, Goose Valley, Shasta County; at altitudes of 6000°—7000° on the Sierra Nevada in Sierra, Madera and Kern Counties, and on Santa Catalina Island; Nevada, on Hunter’s Creek, Washoe County, at an altitude of 6000°.
Occasionally planted as an ornamental plant in the Pacific states, passing into
Sambucus coerulea var. arizonica Sarg.
Sambucus mexicana Sarg., not Presl.
Differing from Sambucus coerulea in its 3—5, usually 3-foliate leaves with usually elliptic long-acuminate leaflets glabrous or slightly pubescent when they appear, 1′—3′ long and ½′—1′ wide, their stipels minute or rudimentary, smaller flower-clusters and fruit not more than ¼′ in diameter.
A tree, often 30° high, with stout spreading branches forming a compact round-topped head, and slender branchlets glabrous or villose pubescent early in the season, usually becoming glabrous. Bark of the trunk about ¼′ thick, the light brown surface tinged with red and broken into long narrow horizontal ridge-like scales. Wood light, soft, close-grained, light brown, with thin lighter-colored sapwood of 2 or 3 layers of annual growth.
Distribution. Banks of streams; Arizona, Grand View Trail, Grand Cañon of the Colorado River and near Flagstaff, Coconino County, Globe, Gila County, and banks of the Rialta near Tucson, Pima County; common; New Mexico, near Silver City, Grant County; southern California (San Diego, Los Angeles, Ventura and Kern Counties).
3. [Sambucus callicarpa] Greene.
Leaves 6′—10′ long, with a stout slightly grooved petiole and 5—7, usually 5, elliptic finely or coarsely serrate leaflets, acuminate and long-pointed at apex, cuneate and often unsymmetric at base, dark green and glabrous on the upper surface, paler and more or less villose-pubescent on the lower surface, especially along the slender midrib, 2½′—5′ long and ½′—2′ wide; petiolules ⅛′—¼′ or that of the terminal leaflet up to 1½′ in length. Flowers on pedicels ⅛′ long, in ovoid to semiorbicular cymes, usually 2½′—3′ long and broad, often somewhat flattened at maturity, on stout peduncles 1½′—3′ in length, about ⅓′ in diameter, with white or yellow slightly obovate petals rounded at apex, and stamens rather shorter than the lobes of the corolla. Fruit about ½′ in diameter, bright red or rarely chestnut color (f. Piperi Sarg.); nutlets smooth.
A tree, occasionally 25°—30° high, with a trunk 10′—12′ in diameter, slender branchlets occasionally puberulous early in the season, becoming glabrous, light brown, separating on the surface into thin scales.
Distribution. River banks in low moist soil, from sea-level in the neighborhood of the coast up to altitudes of 7000°—8000°; coast of Alaska (Skagway), southward along the coast to Marin County, California, and inland to the western slopes of the Cascade and Sierra Nevada Mountains, southward to Amador County; the f. Piperi in western Washington.